Serrognathus (Serrognathus) MOTSCHULSKY, 1861

(G. = Platyprosopus castanicolor)

= Platyprosopus WESTWOOD in HOPE 1845 (G. = Platyprosopus platymelus)


Serrognathus (Serrognathus) bucephalus (PERTY, 1831) [Lucanus, Eurytrachellelus, Dorcus].

Lucanus subcostatus STURM 1826 (numen nudum) [Dorcus] [Java].

Lucanus bubalus PERTY 1831.

Lucanus goliath GISTL. 1831.

Dorcus urus DEJEAN 1837 [nomen nudum] [Java].

Lucanus bilunatus (REINWARDT) DEJEAN 1837 [nomen nudum] [Java].

Dorcus axis DEJEAN 1837 [nomen nudum] [Java].

Dorcus lateralis DEJEAN 1837 [nomen nudum] [Java].

Lucanus punctifrons STURM 1843.

Lucanus (Dorcus) rugifrons HOPE & WESTWOOD 1845 [Java].

Lucanus latus (STURM) GEMMINGER & HAROLD 1868.

Sturm, 1826:165; Perty, 1831:35-36, pl.1; Gistl., 1831:303; Dejean, 1837:193; Sturm, 1843:136; Hope & Westwood, 1845:24; Burmeister, 1847:384, 388; Reiche, 1853:79; Parry, 1864:6, Parry, 1864:48; Vollenhoven, 1865:151; Gemminger & Harold, 1868:956; Parry, 1870:89; Heyne & Taschenberg, 1908:55, pl.8; Berlese, 1910:269; Boileau, 1913:250; Oberthur & Houlbert, 1914:111; Didier, 1931:197; Didier & Sguy, 1952:pl.108; Benesh, 1960:81; Klausnitzer, 1982:46; Maes, 1982:7; Kawano, 1988:8; Maes, 1990:21; Haitlinger, 1991:534; Maes, 1992:87; Haitlinger, 1994:193, 199; Kawano, 1994:9; Mizunuma & Nagai, 1994:271, pl. 107; Kawano, 1997:454.

Java : W. Java (Mt. Gede), E. Java (Mt. Argopuro, Mt. Ijen), Tcibodas, Mt. Salak, Mt. Kawie, Dessa Tjibogo, Pengalengan, Mt. Tjikorai, Mt. Ardjoeno, Toegoe, Bankalan.

Java (4 males, 2 females in col. Ronald Cave, USA).

Natural enemies : Acarina : Canestriniidae : Sandrophela amarae, Sandrophela spectanda (= Canestrinia spectanda), Rugoniphela marloni.

 

Serrognathus (Serrognathus) consentaneus ssp. consentaneus (ALBERS, 1886).

Serrognathus consentaneus (ALBERS, 1886) [Peking].

Dorcus laevidorsis FAIRMAIRE, 1888 [Yunnan].

Dorcus fasolt NOMURA (nec KRIESCHE) 1962, 1970.

Albers, 1886:28; Fairmaire, 1888:15; Kriesche, 1920:118; Nagel, 1926:118; Didier, 1931:203; Benesh, 1960:87; Nomura, 1962:35; Kurosawa, 1970:289, 291; Kurosawa, 1976:8; Kim, 1978:310; Maes, 1982:3; Sugita, 1987:29; Isida & Fujioka, 1988:8; Maes, 1990:21; Maes, 1992:87; Nomura & Lee, 1992:91; Mizunuma & Nagai, 1994:269, pl. 100; Araya, 1996:21.

China : Beijing (Beijing = Peking*)

China : Shandong (Qingdao = Tsingtao = Tsingtau)

China : Jianxi (Jiujiang = Kiukiang)

China : Zhejiang (Ningbo = Ningpo);

North Korea : Pyongyang--3902N-12544N = Pjong-jang;

South Korea : Chemulpo

South Korea : Chosun Peninsula

South Korea : I-Chin-do--3428N-12616E = Jin-do I. (Chollanam-do)

South Korea : Cheju-do I. = Jeju-do I. (Mt. Halla-san--3323N-12630E--Kaewol Bridge);

Japan : Nagasaki Pref. [Tsushima I.--3420N-12920E (Izuhara)].

Note : The type of Dorcus laevidorsis is said to be from Yunnan.

 

Serrognathus (Serrognathus) consentaneus ssp. consentaneus (adapted from Tsukawaki).

Bibliography.

TSUKAWAKI T. (1998) Notes on the genus Dorcus MacLeay, 1819 after the publication of The Lucanid Beetles of the worldin 1994. Gekkan-Mushi, 328:76-84.

 

Serrognathus (Serrognathus) consentaneus ssp. akahorii (TSUKAWAKI, 1998).

Dorcus consentaneus ssp. akahorii TSUKAWAKI, 1998:84, fig. 9 [China : Hubei : Mt. Badong Shan].

China : Hubei : Mt. Badong Shan, 5-VIII-1995 (male HOLOTYPUS in Kanagawa Prefectural Museum of Natural History, Odawara, Japan).

China : Hubei : Mt. Dabie Shan, VI-1992 (1 male PARATYPUS in coll. N. Adachi, K. Fujita, M. Sawai or T. Tsukawaki).

China : Hubei : Mt. Dabie Shan, VI-1996 (2 males PARATYPUS in coll. N. Adachi, K. Fujita, M. Sawai or T. Tsukawaki).

China : Hunan : Mt. Huang Shan, VII-1997 (1 male PARATYPUS in coll. N. Adachi, K. Fujita, M. Sawai or T. Tsukawaki).

China : Sichuan : Mt. Tapa Shan, VII-1992 (11 males PARATYPUS in coll. N. Adachi, K. Fujita, M. Sawai or T. Tsukawaki).

Serrognathus (Serrognathus) consentaneus ssp. akahorii (adapted from Tsukawaki).

Bibliography.

TSUKAWAKI T. (1998) Notes on the genus Dorcus MacLeay, 1819 after the publication of The Lucanid Beetles of the worldin 1994. Gekkan-Mushi, 328:76-84.

 

Serrognathus (Serrognathus) costatus (BOILEAU, 1898) [Eurytrachelus, Eurytrachellelus].

Dorcus metacostatus KIKUTA, 1985 [new name for costatus].

Boileau, 1898:45; Didier, 1931:201; Miwa, 1933:356, pl.2; Didier & Sguy, 1952:pl.107; Benesh, 1960:81; Nomura, 1960:43; Nomura, 1966:69; Maes, 1982:3; Kikuta, 1985:25; Isida & Fujioka, 1988:8; Maes, 1990:21; Araya, Obuchi & Ochi, 1992:32; Maes, 1992:87; Mizunuma & Nagai, 1994:268, pl. 100; Araya, 1996:19, 21.

Japan : Kagoshima Pref. : Amami-oshima I. : Fukumoto

Japan : Kagoshima Pref. : Amami-oshima I. : Kawauchi-gawa near Yuwan

Japan : Kagoshima Pref. : Amami-oshima I. : Mt. Yuwan-dake

Japan : Kagoshima Pref. : Amami-oshima I. : Shinmura

Japan : Kagoshima Pref. : Amami-oshima I. : Mt. Yuidake

Japan : Kagoshima Pref. : Amami-oshima I. : Kinsakubaru

Japan : Kagoshima Pref. : Amami-oshima I. : Gusuku

Japan : Kagoshima Pref. : Amami-oshima I. : Naze

Japan : Kagoshima Pref. Tokunoshima I. : Mt. Amagi-dake

Japan : Kagoshima Pref. Tokunoshima I. : Miky

Japan : Kagoshima Pref. Kakeromajima I.

Japan : Kagoshima Pref. Ukejima I.

 

Serrognathus (Serrognathus) damoiseaui (MAES, 1982).

Eurytrachellelus damoiseaui MAES, 1982:7 [Japan : Clba].

Maes, 1990:21; Tllez, 1990:42; Maes, 1992:87; Araya, 1996:19, 21, 22.

Japan : Clba*, ex col Le Moult (male HOLOTYPUS of Eurytrachellelus damoiseaui MAES, 1982 in col. Institut royal des Sciences naturelles de Belgique ref. I.G.12595).

Serrognathus (Serrognathus) damoiseaui : male HOLOTYPUS from Japan : Cluba

 

Serrognathus (Serrognathus) dorcoides (NAGEL, 1939) [Eurytrachelus, Eurytrachellelus].

Nagel, 1939:329; Benesh, 1960:82; Klausnitzer, 1982:32; Maes, 1982:9; Maes, 1990:21; Maes, 1992:87.

Dorcus titanus ssp. titanus MIZUNUMA & NAGAI (not BOISDUVAL) 1994:pl.104 (part).

Sumatra : S. Sumatra Prov. (W.K. Alahanpandjang*).

 

Serrognathus (Serrognathus) kyanrauensis (MIWA, 1934) [Eurytrachelus].

Miwa, 1934:324, pl.4.

Dorcus nitidus BENESH, (nec KIRSCH) 1950 [Hori, Mizuho, Bukai].

Eurytrachellelus coussementi MAES, 1982:5, 9, 11 [Serrognathus] [cossemati] (male) [Taiwan : Lisan].

Maes, 1990:21; Tllez, 1990:42; Araya, Obuchi & Ochi, 1992:33; Maes, 1992:87; Mizunuma & Nagai, 1994:268-269, pl. 100; Araya, 1996:22.

Taiwan : Taipei Distr. (Mt. Tatun Shan), Taichung Distr. (Lisan), Nantou Distr. (Puli = Hori, Penpuhsi), ? (Mizuho, Bukai).

 

Serrognathus (Serrognathus) platymelus (SAUNDERS, 1854).

Saunders, 1854:46, 50, 52-53, pl.3-4; Motschulsky, 1861:12-13; Vollenhoven, 1861:112, pl.6; Parry, 1864:87; Gemminger & Harold, 1868:957; Harold, 1876:115; Heyden, 1879:338; Lewis, 1883:333; Schonf., 1884:45, 48; Albers, 1886:27; Boileau, 1899:177; Kriesche, 1920:111, 117-119; Kriesche, 1922:124; Didier, 1924; Nagel, 1924:72; Okamoto, 1924:169; Didier, 1925:251; Nagel, 1926:119; Didier, 1927:284; Didier, 1928:53; Nagel, 1930:92; Cho, 1931:58; Didier, 1931:196, 202; Miwa, 1933:356, pl.2; Kriesche, 1935:173; Arrow, 1950:pl.7; Didier & Sguy, 1953:141; Cho, 1957:116; Inahara & Baba, 1959:9; Benesh, 1960:87-88; Nomura, 1960:42-43; Nomura, 1962; Cho, 1963:58; Nomura, 1964:62; Nomura, 1966:68-69; Bomans, 1970:231; Kurosawa, 1970:289; Ratcliffe, 1970:256; Kurosawa, 1976; Kim, 1978:309; Nakane, 1979:1; Klausnitzer, 1982:46; Maes, 1982:3, 5; Fujita & Ichikawa, 1985:9-10, 13; Kurosawa, 1985:343; Lee, Kim & Kim, 1985:417; Fujita & Ichikawa, 1986:28-29; Nakane, 1987:135; Nameta, 1987:40; Sugita, 1987:29; Isida & Fujioka, 1988:8-9; Araya, 1989:7; Bomans, 1989:8; Fujita & Okuda, 1989:24-26; Sato & Saito, 1989:300; Maes, 1990:21; Bomans, 1991: 149; Araya, 1992:61-66; Araya & Obuchi, 1992:33, 37; Araya, Obuchi & Ochi, 1992:32; Nomura & Lee, 1992:91; Maes, 1992:88; Haitlinger, 1993:344; Ma, 1993:278-279; Haitlinger, 1994:193; Adachi, 1994:20; Mizunuma & Nagai, 1994:260, 269-270, pl. 101-103; Araya, Oobuchi & Ochi, 1995:12; Araya, 1996:18-19, 21; Kawano, 1997:454; Bomans, Drumont & Cammaerts, 1997:587-589.

 

Serrognathus (Serrognathus) platymelus ssp. castanicolor MOTSCHULSKY, 1861 [Dorcus].

Eurytrachelus titanus subsp. fasolt KRIESCHE 1920 [Korea : Pjong-jang].

N. China;

North Korea : Pyongyang--3902N-12544N = Pjong-jang;

South Korea : Chosun Peninsula, I-Chin-do--3428N-12616E = Jin-do I. (Chollanam-do), Cheju-do I. = Jeju-do I. (Mt. Halla-san--3323N-12630E--Kaewol Bridge);

Korea : Kyonggi-do (Kwangnung);

Japan : Nagasaki Pref. [Tsushima I.*--3420N-12920E (Seto), Goto I.].

 

Serrognathus (Serrognathus) platymelus ssp. pilifer (VOLLENHOVEN, 1861).

Eurytrachelus pilifer VOLLENHOVEN, 1861 [Japan].

Serrognathus titanus BOISDUVAL; Ratcliffe, 1970.

Dorcus pilifer (VOLLENHOVEN); Araya, 2000:11-12.

Dorcus titanus pilifer (VOLLENHOVEN); Araya, 2001:272.

Dorcus titanus pilifer (VOLLENHOVEN); Hosoya, Honda & Araya, 2001:344.

        Japan : Honshu I. : Niigata Pref.

        Japan : Honshu I. : Fukui Pref. (citated by Araya, 2001).

        Japan : Honshu I. : Izu Is. : Ooshima I., 3446N-13924E

        Japan : Honshu I. : Izu Is. : Toshima I.

        Japan : Honshu I. : Izu Is. : Niijima I., 3424N-13917E

        Japan : Honshu I. : Izu Is. : Shikinejima I.

        Japan : Honshu I. : Izu Is. : Miyakejima I., 3404N-13930E

        Japan : Honshu I. : Kanagawa Pref. : Mutsu-ura-cho, 3519N-13937E

        Japan : Honshu I. : Kanagawa Pref. : Sarushima I., 3516N-13941E

        Japan : Honshu I. : Kyoto Pref. : Kyoto, 3505N-13551E : Sakyo : Kyoto University (1 ex. citated by Hosoya, Honda & Araya, 2001).

        Japan : Honshu I. : Nara Pref. : Mt. Odaigahara, 3415N-13607E

        Japan : Honshu I. : Hyogo Pref. : Takarazuka

        Japan : Honshu I. : Hyogo Pref. : Sonoda

        Japan : Honshu I. : Tottori Pref. : Oki Is. : Togo I.

        Japan : Shikoku I.

        Japan : Kyushu I. : Nagasaki Pref. : Nagasaki, 3247N-12954E

        Japan : Kyushu I. : Goto Is. : Fukuejima I., 3241N-12844E

        Japan : Kyushu I. : Miyazaki Pref. : Tano

        Japan : Kagoshima Pref. : Tanegashima I., 3034N-13059E

        Japan : Kagoshima Pref. : Yakushima I., 3022N-13031E : Miyanoura

        Japan : Kagoshima Pref. : Kumage Is. : Takeshima I.

        Japan, col. Von Siebold (male LECTOTYPUS, male PARALECTOTYPUS of Eurytrachelus pilifer VOLLENHOVEN, 1861 in Rijksmuseum, Leiden).

        Japan, col. Burger (female PARALECTOTYPUS of Eurytrachelus pilifer VOLLENHOVEN, 1861 in Rijksmuseum, Leiden).

 

Serrognathus platymelus ssp. pilifer : LECTOTYPUS and PARALECTOTYPUS OF Eurytrachelus pilifer VOLLENHOVEN (adapted from Araya, 2000)

Bibliography.

ARAYA K. (2000) An account of a visit to European Museums. 2. The lucanid specimens in the van Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke Historie, Leiden. Gekkan-Mushi, 350:4-16.

ARAYA K. (2001) New records of Ceruchus lignarius monticola Nakane, 1978 (Coleoptera, Lucanidae) from Fukui Prefecture, Central Japan, with reference to Lucanid fauna of Fukui Prefecture. Sukunahikona, Special Publication of the Japan Coleopterological Society, 1:271-273.

HOSOYA T., HONDA M. & ARAYA K. (2001) Genetic variation of 16S rRNA gene observed in Ceruchus lignarius and Dorcus rectus rectus (Coleoptera : Lucanidae). Entomological Science, 4(3):335-344.

 

Serrognathus (Serrognathus) platymelus ssp. hachijoensis FUJITA & OKUDA, 1989.

Japan : Tokyo Met. [Izu Is. (Hachijojima I.--3307N-13948E {Sueyoshi*, Kashidate*})].

 

Serrognathus (Serrognathus) platymelus ssp. takaraensis FUJITA & ICHIKAWA, 1985.

Japan : Kagoshima Pref. [Tokara Is. (Takarajima I.*)].

 

Serrognathus (Serrognathus) platymelus ssp. elegans (BOILEAU, 1899) [Eurytrachelus, Eurytrachellelus].

Japan : Kagoshima Pref. [Amami-oshima I.* (Yamamoto, Kawauchi-gawa near Yuwan, Mt. Yuwan-dake, Kinsakubaru)].

 

Serrognathus (Serrognathus) platymelus ssp. tokunoshimaensis FUJITA & ICHIKAWA, 1985.

Japan : Kagoshima Pref. [Tokunoshima I. (Miky*, Isen-ch*, Hetono*, Gozend, Mt. Amagi-dake, Itokina, Yonama), Yorojima I.*, Ukejima I.].

 

Serrognathus (Serrognathus) platymelus ssp. okinoerabuensis FUJITA & ICHIKAWA, 1985.

Japan : Kagoshima Pref. [Okinoerabujima I.* (Mt. Ohyama*)].

 

Serrognathus (Serrognathus) platymelus ssp. okinawanus (KRIESCHE, 1922) [Eurytrachelus].

Japan : Okinawa Pref. [Okinawa I.*--2630N-128E = Okinawahonto I. (Yona, Izumi--Motobu-ch, Mt. Yonahadake, Kunigami-son), Kumejima I., Tokashikijima I., Minnajima I.--2643N-12747E, Iheyajima I., Iejima I., Kourijima I.].

 

Serrognathus (Serrognathus) platymelus ssp. daitoensis FUJITA & ICHIKAWA, 1986.

Japan : Okinawa Pref. [Kitadaitjima I.*, Minamidaitjima I. (Dait-jinja*)].

 

Serrognathus (Serrognathus) platymelus ssp. sakishimanus NOMURA, 1964.

Eurytrachelus platymelus MIWA (nec SAUNDERS) 1933.

Dorcus titanus okinawanus NOMURA (nec KRIESCHE) 1962.

Japan : Okinawa Pref. [Ishigakijima I.*--2424N-12410E (Mt. Omoto* = Mt. Omotodake, Mt. Banna* = Mt. Bannadake), Iriomotejima I.*--2420N-12346E (Mt. Hateruma*, Shirahama), Yonagunijima I.--2428N-12257E (Mt. Urabe*, Sonae*, Higawa), Taketomijima I.].

 

Serrognathus (Serrognathus) platymelus ssp. platymelus (SAUNDERS, 1854) [Platyprosapus, Eurytrachellelus, Eurytrachellus] [platymerus].

Dorcus lateralis SAUNDERS 1854 [lapsus for lateralis DEJEAN from Java].

Dorcus obscurus SAUNDERS 1854 [China].

Dorcus marginalis SAUNDERS 1854 [China].

Eurytrachelus titanus subsp. fafner KRIESCHE 1920 Tonkin : Chiem-hoa, Mauson Mts.].

Eurytrachelus titanus subsp. hangul KRIESCHE 1920 [Siao-Lou].

Eurytrachelus platymelus var. typhoniformis NAGEL 1924 [Serrognathus] [Yunan].

Eurytrachelus prometheus DIDIER 1925 [Serrognathus] [China : Kweitscheou].

Eurytrachelus hymir KRIESCHE 1935 [Serrognathus] [China : Ssetschuan : Mts. near Kwanhsien].

Dorcus titanus ssp. westermanni MIZUNUMA & NAGAI (not HOPE) 1994.

Dorcus titanus BOISDUVAL; KAWANO 1997.

Serrognathus titanus platymelus f. dissimilis BOMANS, DRUMONT & CAMMAERTS 1997 [China : Sichuan : You-Yang].

Myanmar = Burma : Kachin State (Putao District--Sumpra Bum), Shan State (Muang Yoo near Mong Yang);

China* : Canton, Sichuan = Szetchuan (Siao-lou-lou-chan, Kwanhsien, Mt. Kinfuschan, Mt. Yunling, Mt. Emei Shan, You-Yang), Hubei (Mt. Wufeng Shan), Hunan (Wuling Mts.), Yunnan (Jinghong, Nyenhangli), Zhejiang (Wenzhou, Mt. Siming Shan), Fujian = Fokien (At Chong, Kuatun), Guangdong = Kouantoung (Tscha-jiu-san, Fung-wan, Tong-kung-san, Yu-sa-tan), Hainan I.;

Thailand : Chiang Rai (Mae Jan, Chiang Kong), Chiang Mai (Mt. Doi Pui, Chiang Dao--1915N, Chiang Mai--1850N-9857E, Samoeng, Sansai, Mt. Doi Saket, Mt. Doi Suthep, Maetaeng, Maesa, Fang), Tak (Umphang), Petchaburi (Khao Khor);

Laos: Thakek, Xieng-Khouang, Parkading, P.K. Khouai--Vientiane, Ban Houei Sai, Louang Prabang, Muong Soui, Ban Sam Thong, Ban Van Heua, Paksong, Sayabouri, Thonpheng, Samnea;

Vietnam : Annam, Tonkin (Mt. Mauson, Chiem-hoa, Sia-Lou), Banmethuot, Tam Dao near Vinh Yen.

Saproxylophagous.

Natural enemies : Acarina : Canestriniidae : Cooremania wauensis, Sandrophela rodioni.

Note : The synonyms given in this subspecies are taken from titanus in Benesh (1960) for the distribution area of platymelus. It's possible that various of them are in reality good species or subspecies.

 

Serrognathus (Serrognathus) sika (KRIESCHE, 1920) [Eurytrachelus, Eurytrachellelus].

Eurytrachelus creus DIDIER 1927 [Formose : Taipeli].

Eurytrachellelus coussementi MAES 1982 [Serrognathus] (female) [Lisan].

Kriesche, 1920:118; Didier, 1927:284-285; Didier & Sguy, 1952:pl.102; Benesh, 1960:87; Maes, 1982:7; Maes, 1990:21; Maes, 1992:89; Sakaino, Yu & Chu, 1994:123; Mizunuma & Nagai, 1994:269, pl. 102; Araya, 1996:21.

Taiwan : Taoyuan Distr. (Mt. Rara Shan), Taipei Distr. (Taipei), Taichung Distr. (Lisan), Hualien Distr. (Taihorin*), Kaohsiung Distr. (Liukuei), ? (Kosempo*, Polischa*, Fuhoscho*, Hoozan*, Teraso, Taihorinsho*, Kankau--Koshun), Lutao I.--2239N-12128E.

 

Serrognathus (Serrognathus) titanus (BOISDUVAL, 1835) [Lucanus].

Boisduval, 1835:237, pl.6; Hope & Westwood, 1845:20; Burmeister, 1847:384; Albers, 1893:72; Heyne & Taschenberg, 1908:55; Boileau, 1913:249; Gravely, 1915:426; Didier, 1915 (?1925):426; Didier, 1931:199; Gardner, 1935:8, pl.1; Arrow, 1943:137; Arrow, 1950:104, pl.7; Didier & Sguy, 1952:pl.101; Didier & Sguy, 1953:52; Sguy, 1954:88; Benesh, 1960:86; De Lisle, 1977:491; Klausnitzer, 1982:14; Maes, 1982:9; Lacroix, 1983:17; Weng-Yew Tung, 1983:47, lam4; Fujita & Ichikawa, 1985:9; Kawano, 1988:22; Kawano, 1989:17; Lacroix, 1990:14; Maes, 1990:21; Araya, Obuchi & Ochi, 1991:23; Bomans, 1991:30; Maes, 1992:89; Kawano, 1992:12; Kawano, 1993:13; Mizunuma & Nagai, 1994:270-271, pl. 104-107; Araya, 1994:3.

 

Serrognathus (Serrognathus) titanus ssp. titanus (BOISDUVAL, 1835).

Lucanus titanus BOISDUVAL, 1835 [Celebes : Manado].

Lucanus briareus HOPE & WESTWOOD, 1845.

Dorcus Titan BOISDUVAL; Ritsema, 1891:238.

Prosopocoelus lumawigi DE LISLE, 1977 (female).

      Malaya = Malacca : Pinang : Pinang I. = Penang, 522N-10012E

      Malaya = Malacca : Perak : Cameron Highlands : Taphh

      Malaya = Malacca : Perak : Kampong Sahom

      Malaya = Malacca : Perak : Taiping, 451N-10041E

      Malaya = Malacca : Pahang : Cameron Highlands

      Malaya = Malacca : Singapore

      Borneo : Sarawak

      Borneo : Sabah : Crocker Range

      Borneo : Sabah : Ranau, 20-IV-2001, leg. Chew (1 male in col. Ronald Cave, USA).

      Borneo : Sabah : Ranau, IV-2001, leg. Chew (1 male in col. Ronald Cave, USA).

      Borneo : Sabah : Trus Madi, IV-2001, leg. Chew (2 males in col. Ronald Cave, USA).

      Borneo : W. Kalimantan : Singkawang : Mt. Bawang

      Borneo : W. Kalimantan : Balaikarangan

      Borneo : E. Kalimantan : Balikpapan

      Borneo : S. Kalimantan : Bandjermasin

      Sumatra : N. Sumatra : Berastagi

      Sumatra : Nias I.

      Sumatra : W. Sumatra Prov. : Fort de Kock

      Sumatra : W. Sumatra Prov. : Muaralabuh

      Sumatra : W. Sumatra Prov. : Padang

      Sumatra : Lampung Prov.

      Sumatra : ? : Ajer-Mantcior

      Sumatra : ? : Mt. Liang Gagang

      Sumatra : ? : Mt. Dadap

      Sumatra : ? : Tambang Salida

      Sumatra : ? : Ampugadang

      Sulawesi = Celebes : Minahassa Peninsula : Manado, 131N-12447E (TYPUS of Lucanus titanus BOISDUVAL, 1835).

      Sulawesi = Celebes : Minahassa Peninsula : Tondano, 114N-12451E)

      Sulawesi = Celebes : Bantimulung

      Sulawesi = Celebes : Palu, 046S-11949E : Palolo

      Sulawesi = Celebes : Sinjai

      Sulawesi = Celebes : Bantimurung

      Sulawesi = Celebes : Banggai Is. : Peleng I., 118S-12305E

      Philippines : Luzon I. : Mountain Province, II-1988 (1 male in coll. Mizunuma, Japan).

      Philippines : Luzon I. : Mountain Province, 13-VIII-1988 (1 male in coll. Mizunuma, Japan).

      Philippines : Marinduque I., 1324N-12157E, VI-1991 (1 male in coll. Mizunuma, Japan).

      Philippines : Samar I., 1149N-12514E, IX-1991 (1 male in coll. Mizunuma, Japan).

      Philippines : Leyte I., 1054N-12454E, XII-1991 (1 male in coll. Mizunuma, Japan).

      Philippines : Mindanao I. : Mt. Kintanlad, III/IV-1990 (1 female in coll. Mizunuma, Japan).

      Philippines : Mindanao I. : Mt. Apo, 708N-12509E, 8-III-1977 (1 male in coll. Mizunuma, Japan).

      Philippines : Mindanao I. : Mt. Apo, 708N-12509E, III-1984 (1 male in coll. Mizunuma, Japan).

      Philippines : Mindanao I. : Mt. Apo, 708N-12509E, III-1986 (1 male in coll. Mizunuma, Japan).

      Philippines : Mindanao I. : Surigao, 1985 (1 male in coll. Mizunuma, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

RITSEMA C. (1891) Note XXXV: Two new species of the lucanoid genus Cyclommatus Parry. Notes Leyden Museum, XIII:235-238.

 

Serrognathus (Serrognathus) titanus ssp. palawanicus LACROIX, 1983.

Serrognathus titanus ssp. palawanicus LACROIX, 1983 (Palawan).

Serrognathus titanus ssp. palawanicus LACROIX; Bomans & Benoit, 2005:298.

      Philippines: Palawan I. : Pinigisan

      Philippines: Palawan I. : Brooke's Point, 854N-11800E, IX-1983 (1 male in coll. Mizunuma, Japan).

      Philippines: Palawan I. : Brooke's Point, 854N-11800E, VI-1987 (2 males, 1 female in coll. Mizunuma, Japan).

      Philippines: Palawan I. : Brooke's Point, 854N-11800E, IX-1988 (1 male in coll. Mizunuma, Japan).

      Philippines: Palawan I., VIII-1971, leg. Monguillon (Male and female TYPUS in MNHN, Paris).

Bibliography.

BOMANS H.E. & BENOIT P. (2005) Inventaire provisoire comment des Coloptres Lucanides de lArchipel des Philippines (98eme Contribution a ltude des Coloptres Lucanides). Lambillionea, CV(2):283-301.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Serrognathus (Serrognathus) typhon (BOILEAU, 1905) [Eurytrachelus, Eurytrachellus, Eurytrachellelus].

Boileau, 1905:17; Didier, 1931:200; Sguy, 1954:88; Benesh, 1960:87; Maes, 1982:9; Maes, 1990:21; Maes, 1992:89; Mizunuma & Nagai, 1994:pl. 105-106.

Dorcus titanus ssp. titanus MIZUNUMA & NAGAI (not BOISDUVAL) 1994:pl.105-106 (in part).

Sulawesi = Celebes;

Philippines : Luzon I., Mindanao I.

Unknown locality (labelled as Celebes : Arfak Is.), I-1979, leg. E. Coussement (1 male in Museo Entomolgico de Len).

Serrognathus typhon : unknown locality, probably Sulawesi (labelled as Celebes : Arkak Is.) in col. Museo Entomolgico de Len

 

Serrognathus (Serrognathus) westermanni (HOPE, 1842) [Dorcas, Eurytrachellelus, Dorcus].

Hope, 1842:127; Hope, 1843:347; Hope, 1843:106; Didier, 1931:204; Benesh, 1960:86; Maes, 1982:5; Maes, 1990:21; Maes, 1992:89; Mizunuma & Nagai, 1994:269, pl. 101.

India : Sikkim, West Bengal (Mangpu, Darjeeling, Pashok, Dalfa Hills, Kurseong), Assam (Sibsagar), Meghalaya (Cherrapunji, Khasi Hills, Shillong);

Bhutan : Pedong;

Bangladesh : Sylhet*.


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