Calcodes WESTWOOD, 1834
(G. = Lucanus aeratus)
= Chalcodes
BURMEISTER, 1847 (G. = Lucanus aeratus)
= Odontotyranus
SEGUY, 1955 (G. = Odontolabis striatus)
Calcodes
aeratus (HOPE, 1834).
Lucanus
aeratus HOPE, 1834:118 [Tenasserim] [Anoplocnemus, Odontolabis, Chalcodes]
[aerata].
Hope,
1835:99, pl.14; Castelnau, 1840:172; Hope & Westwood, 1845:16; Burmeister,
1847:361; Westwood, 1848:22, pl.10; Parry, 1864:pl.7; Parry, 1870:74; Leuthner,
1885:473, pl.97; Zang, 1905:214; Heyne & Taschenberg, 1908:52; Gravely,
1915:416; Arrow, 1937:241, pl.3; Arrow, 1950:209, pl.20; Arrow, 1951:pl.15;
Didier & Séguy, 1952:pl.41; Didier & Séguy, 1953:91; Benesh, 1960:121;
Weinreich, 1972; Lacroix, 1978:262; Lacroix, Ratti & Taroni, 1983:2;
Weng-Yew Tung, 1983:49, pl.4; Lacroix, 1984:34-35, pl.1; Kikuta, 1986:135;
Nagai, 1986:3, 10; Bartolozzi & Bomans, 1990:438; Bomans, 1991:145; Maes,
1992:22.
Calcodes
aeratus (HOPE); Mizunuma & Nagai, 1994:16-17, fig. 38(a-f), p. 230, pl. 38.
Araya,
1996:17.
Myanmar =
Burma : Tenasserim* State.
Thailand
: Surat Thani.
Malaya = Malacca : Kedah.
Malaya : Pinang : Pinang I., 5°22N-100°12E
(= Prince of Wales I.).
Malaya : Perak : Taiping, 4°51N-100°41E.
Malaya : Perak : Kampong Sahom.
Malaya : Perak : Cameron Highlands : Taphh.
Malaya :
Pahang : Cameron Highlands.
Malaya :
Pahang : Tengah-Geb.
Malaya :
Pahang : Fraser's Hills.
Malaya : Singapore : Chapais.
Borneo : Natuna Is. : Bunguran Besar I.,
3°56N-108°15E.
Borneo : W. Kalimantan : Sambas.
Sumatra : W. Sumatra Prov. : Sinkep I.
Sumatra : S. Sumatra Prov. : Palembang.
Sumatra : Siboga.
Java.
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Calcodes
baderi (BOMANS & BARTOLOZZI, 1990) [Chalcodes, Odontolabis].
Bomans
& Bartolozzi, 1990:433; Maes, 1992:22; Mizunuma & Nagai, 1994:16, 231, pl.
39; Ipsen, 1995:1; Ikeda, 1996:4.
Sulawesi = Celebes* : Palopo--2°55S-120°06E
(Pulu Pulu).
Calcodes carinatus (LINNAEUS, 1758) [Scarabaeus, Odontolabis,
Chalcodes] [carinata] [India].
= Lucanus bengalensis TENNENT 1859.
= Odontolabis cingalensis PARRY 1864
[Ins. Taprobana].
= Odontolabis intermedia DEYROLLE
1864 [Ceylon].
= Odontolabis
nigrita DEYROLLE 1864 [Chalcodes] [nigritus] [Ceylon].
= Chalcodes
srilankaensis BOMANS 1974 [Ceylan].
Linnaeus,
1758:353; Linnaeus, 1764:34; Linnaeus (ed. Gmelin), 1790:1590; Herbst,
1790:315; Muller, 1776:97; Thunberg, 1806:193, pl.12; Schonherr, 1817:323;
Tennent, 1859:276; Deyrolle, 1864:314; Parry, 1864:16, 77, pl.10; Sharp,
1884:18; Leuthner, 1885:474, pl.97; Heyne & Taschenberg, 1908:52, pl.7;
Boileau, 1913:243; Gravely, 1915:416; Arrow, 1937:241, pl.1; Arrow, 1950:207,
pl.20; Arrow, 1951:pl.15; Didier & Séguy, 1952:pl.40; Didier & Séguy,
1953:91; Benesh, 1960:114; Bomans, 1974:181; Lacroix, 1978:258; Numhauser,
1981:74; Lacroix, 1984:39-43, pl.3-4;
Bartolozzi & Bomans, 1988:88-89; Bartolozzi & Bomans, 1990:433; Maes,
1992:22; Mizunuma & Nagai, 1994:229, pl. 38.
Sri Lanka
= Ceylon : Western Prov. [Colombo Distr.
(Labugama), Kalutara Distr. (Agalawatta)], Sabaragamuwa Prov. [Ratnapura Distr.
(Rakwana, Bulutota, Gilimale, Sinharaja Jungle 3 mi S Weddagala)], Central
Prov. [Kandy Distr. (Kandy, Karabagala--Willomalai Jungle), Nuwara Eliya Distr.
(Bogawantalawa, Maskeliya, Lindula)], Uva Prov. [Badulla Distr. (Ohiya--W.
Haputale, Haldumulla)], Southern Prov. [Galle Distr. (Udugama--Kaneliya,
Kaneliya Jungle, Sinharaja Jungle, Galle)], ? [Taprobana, Mousakande].
Calcodes cephalotes (LEUTHNER, 1885) [Odontolabis, Chalcodes].
= Odontolabis striata WESTWOOD (nec
DEYROLLE) 1874 (en parte).
Westwood, 1874; Leuthner, 1885:478, pl.97;
Benesh, 1960:115; De Lisle, 1976:276; Klopp, 1977:60; Lacroix, 1978:259;
Lacroix, 1984:45-47, pl.5; Nagai, 1986:8; Bartolozzi & Bomans, 1990:438;
Maes, 1992:22.
Borneo*.
Calcodes cupreiventris (MOLLENKAMP, 1900).
Odontolabis cupreiventris MOLLENKAMP, 1900:45 [Borneo].
Mollenkamp, 1903:348; Didier & Séguy,
1953:98; Benesh, 1960:121; Lacroix, 1978:262; Lacroix, 1984:36-37, 46, pl.5;
Nagai, 1986:10; Bartolozzi & Bomans, 1990:438; Maes, 1992:22.
Calcodes
cupreiventris (MOLLENKAMP); Mizunuma & Nagai, 1994:17, fig. 39(a-b, f), p. 230,
pl. 38.
Araya,
1996:17.
Borneo :
Sarawak : Mt. Serapi near Kuching.
Borneo :
W. Kalimantan : Mt. Bawang.
Sumatra : Jambi Prov. : O.K.Siantar-Simpang
Raja.
Sumatra : Sibolangit.
Sumatra : Langkat-Balei Gadjah.
Sumatra : Deli.
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Calcodes invitabilis (MIZUNUMA, 1994).
Odontolabis invitabilis MIZUNUMA in MIZUNUMA & NAGAI, 1994:16-17, fig.
37(a-c). p. 229, 230, pl. 139, fig. 183(1-2) [Berastagi].
Sumatra : N. Sumatra Prov. : near Berastagi,
IV-1993 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan).
Sumatra : N. Sumatra Prov. : near Berastagi,
V/VII-1993 (3 males PARATYPUS in col. T. Mizunuma, Japan) (1 male PARATYPUS in
col. H. Fujita, Japan).
Sumatra : N. Sumatra Prov. : near Berastagi
(1 male PARATYPUS in col. H. Yasuoka, Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Calcodes
kazuhisai (MIZUNUMA, 1994).
Odontolabis
kazuhisai MIZUNUMA in MIZUNUMA & NAGAI, 1994:16-17, fig. 36(a-b, f), pl. 38,
fig. 184(1-3) [ Mt. Singgalang].
Kawano, 1997:454.
Sumatra : W. Sumatra Prov. : Padang :
Padangpanjang : Mt. Singgalang, 7-VIII-92, leg. Kazuhisa Fujita (male
HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (1 male PARATYPUS in col. T. Mizunuma, Japan).
Sumatra : W. Sumatra Prov. : Padang :
Padangpanjang : Mt. Singgalang, 28-VII-92, leg. Kazuhisa Fujita (female
ALLOTYPUS in Kanagawa Prefectural Museum, Japan).
Sumatra : W. Sumatra Prov. : Padang :
Padangpanjang : Mt. Singgalang, II/III-93 (16 males PARATYPUS in col. T.
Mizunuma, Japan).
Sumatra : W. Sumatra Prov. : Padang :
Padangpanjang : Mt. Singgalang, III-93 (4 males PARATYPUS, 1 female PARATYPUS
in col. H. Fujita, Japan).
Sumatra : W. Sumatra Prov. : Padang :
Padangpanjang : Mt. Singgalang, V-93 (30 males PARATYPUS, 30 females PARATYPUS
in col. T. Mizunuma, Japan) (2 females PARATYPUS in col. T. Mizukami, Japan).
Sumatra : W. Sumatra Prov. : Padang :
Padangpanjang : Mt. Singgalang (6 males PARATYPUS, 2 females PARATYPUS in col.
H. Yasuoka, Japan) (4 males PARATYPUS, 2 females PARATYPUS in col. M. Suzumura,
Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Calcodes
lecourti (BOMANS, 1997) [Chalcodes].
Bomans,
1997 : 29-30.
Sulawesi
= Celebes : Palopo--2°55S-120°06E (Rantepao--Pulu-Pulu*).
Calcodes
martinii (IPSEN, 1995) [Chalcodes].
Ipsen,
1995:1-5.
Sulawesi
: Palopo--2°55S-120°06E (Rantepao*).
Calcodes
nagaii (IKEDA, 1996) [Odontolabis].
Ikeda, 1996:3-6.
Sulawesi = Celebes : Palopo--2°55S-120°06E
(Pulu Pulu*).
Calcodes nishiyamai (MIZUNUMA, 1994).
Odontolabis nishiyamai MIZUNUMA in MIZUNUMA & NAGAI, 1994:16-17, fig.
40(a-h), pp. 231-232, pl. 39, fig. 191(1-14) [Ende].
Ikeda,
1996:3-4.
Lesser
Sunda Is. : Flores I. (8°37S-121°19E) : near Ende, XI/XII-1991 (male HOLOTYPUS,
3 males PARATYPUS in Kanagawa Prefectural Museum, Japan) (female ALLOTYPUS, 84
males PARATYPUS in col. T. Mizunuma,
Japan) (4 males PARATYPUS in col. M. Suzumurai, Japan) (10 males PARATYPUS in
col. H. Fujita, Japan) (6 males PARATYPUS, 3 females PARATYPUS in col. M. Baba,
Japan) (3 males PARATYPUS in col. T. Mizukami, Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Calcodes
ritsemae (BOILEAU, 1899) [Odontolabis, Chalcodes] [ritsemai].
= Odontolabis
liewi DE LISLE 1976 [Malaya : Maxwell's Hill].
= Odontolabis
albolateralis KLOPP 1977 [Malaya : Perak : Taiping].
Boileau,
1899:42; Benesh, 1960:119; De Lisle, 1976:274-277; Klopp, 1977:60; Lacroix,
1984:48-49, pl.6; Bartolozzi & Bomans, 1990:438; Maes, 1992:23; Mizunuma
& Nagai, 1994:229, pl. 38.
Malaya = Malacca*
: Perak (Taiping--4°51N-100°41E, Taiping--Maxwell's Hill), Pahang (Cameron
Highlands);
Sumatra : N. Sumatra Prov. (Dolok Merangir).
Calcodes rorekaotimbuensis (NAGAI & ISOGAI, 1987) [Odontolabis, Chalcodes].
Nagai & Isogai, 1987:9-11; Bartolozzi
& Bomans, 1990:436; Maes, 1992:23; Mizunuma & Nagai, 1994:16, 231, pl.
39; Ipsen, 1995:1; Ikeda, 1996:3-4.
Sulawesi = Celebes : Mt.
Rorekaotimbu--Puncak Dingin*.
Calcodes
rubens (LACROIX, 1978) [Chalcodes].
Lacroix,
1978:264; Lacroix, 1984:37-38, 42, pl.4; Bartolozzi & Bomans, 1990:438;
Maes, 1992:23; Mizunuma & Nagai, 1994:230.
Malaya :
Pahang (Kuala-Lipis*);
Borneo :
Sarawak (Mt. Serapi near Kuching), W. Kalimantan (Mt. Bawang near Singkawang).
Calcodes
sarasinorum (HELLER, 1898) [Odontolabis, Chalcodes].
Heller,
1898:21, pl.3; Benesh, 1960:119; De Lisle, 1976:277; Lacroix, 1984:49-50, pl.6;
Nagai & Isogai, 1987:9-10; Bartolozzi & Bomans, 1990:438Maes, 1992:27;
Mizunuma & Nagai, 1994:16, 231, pl. 39; Ikeda, 1996:4.
S.
Sulawesi = Celebes : Mt. Bonthain*
S.
Sulawesi : Sinjai
S.
Sulawesi : Mt. Bawakaraeng, I-2001, leg. M. Djufri (2 males, 1 female in col.
Ronald Cave, USA).
Calcodes
striatus (DEYROLLE, 1864) [Odontolabis, Chalcodes,
Odontotyranus] [striata].
Deyrolle,
1864; Westwood, 1874:360, pl.3; Boileau, 1913:241; Didier & Séguy,
1952:pl.39; Séguy, 1955:35; Benesh, 1960:120; De Lisle, 1976:276; Klopp,
1977:60; Klausnitzer, 1982:29; Lacroix, 1984:44-45, pl.5; Nagai, 1986:3, 8-10;
Bartolozzi & Bomans, 1990:438; Maes, 1992:23; Mizunuma & Nagai,
1994:229, pl. 38, 139; Araya, 1994:4.
Malaya = Malacca* : Johor
(Kuantan--3°50N-103°18E);
Borneo :
Sarawak (Mt. Serapi near Kuching), Sabah (Telupid, Crocker Range, Sandakan), W.
Kalimantan (Pontianak, Putusibau, Karimata I.), ? (Gunong-Gadin-Lundu);
Sumatra : W. Sumatra Prov. (Mt. Singgalang),
Belitung I.;
Java.
Note :
Perhaps it's safe to use the subgenus Odontotyranus SEGUY for the Chalcodes
with spinose jugae, like C. striatus, C. ritsemae and C.
cephalotes.
Bibliography.
ARAYA
K. (2000)
An account of a visit to European Museums. 2. The lucanid specimens in the van
Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke
Historie, Leiden. Gekkan-Mushi, 350:4-16.
Calcodes
taronii (BOMANS & BARTOLOZZI, 1990) [Chalcodes, Odontolabis] [tarorii].
Bomans
& Bartolozzi, 1990:436; Maes, 1992:23; Mizunuma & Nagai, 1994:16, 231,
pl. 39; Ipsen, 1995:1; Ikeda, 1996:3-4.
S. Sulawesi = Celebes* :
Palopo--2°55S-120°06E (Puncak*, Pulpul*, Pulu Pulu), Bantaeng*.