Lamprima LATREILLE, 1806 (G

Odontolabis HOPE, 1842

(G. = Odontolabis cuvera)

= Anoplocnemus HOPE 1843 (G. = Lucanus burmeisteri)

= Odontolabis HOPE & WESTWOOD 1845 (G. = Lucanus delesserti)

= Anoplocnemus BURMEISTER 1847 (G. = Lucanus alces)

Odontolabis alces (FABRICIUS, 1775).

Lucanus alces FABRICIUS, 1775:1 (Asia).

Fabricius, 1781:1

Lucanus lama OLIVIER, 1789:8-9, 14-15, pl.2-3 [Indes Orientales].

Herbst, 1790:297; Linnaeus (Ed. Gmelin), 1790:1588; Fabricius, 1792:236, 238; Olivier (ed. Illiger), 1800:54, 62; Fabricius, 1801:248, 250; Olivier (ed. Sturm), 1802:1, 3, pl.2, 4; Thunberg, 1806:104, 188; Schonherr, 1817:324

Lucanus dux WESTWOOD, 1842:124 [Manilla].

Lucanus cumingii HOPE & WESTWOOD, 1845:17 [Manilla].

Burmeister, 1847:357; Westwood, 1848:17, pl.8; Parry, 1863:453; Leuthner, 1885:443, pl.89; Heyne & Taschenberg, 1908:51; Boileau, 1913:238; Didier, 1929:82; Didier & Séguy, 1952:223; Didier & Séguy, 1952:pl.42; Didier & Séguy, 1953:17, 33; Séguy, 1954:87; Benesh, 1960:112; De Lisle, 1976:276; Numhauser, 1981:74; Klausnitzer, 1982:42; Lacroix, 1984:63-65, pl.8; Maes, 1992:23; Kawano, 1993:12

Odontolabis alces (FABRICIUS); Mizunuma & Nagai, 1994:228, pl. 35, fig.175(1-11).

Kawano, 1997:454.

Odontolabis alces (FABRICIUS); Bomans & Benoit, 2005:294.

Ø Philippines : Luzon I. : Mt. Banahao

Ø Philippines : Luzon I. : Manilla, 14°35N-120°57E

Ø Philippines : Luzon I. : Banaue, IV-1990 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Luzon I. : Quezon Prov. : Baler

Ø Philippines : Mindoro I., 12°58N-121°03E

Ø Philippines : Marinduque I., 13°24N-121°57E, II-1988 (2 males in coll. Mizunuma, Japan).

Ø Philippines : Catanduanes I., 13°46N-124°16E

Ø Philippines : Samar I., 11°49N-125°14E, III-1990 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Samar I., 11°49N-125°14E, VI-1990 (1 male, 1 female in coll. Mizunuma, Japan).

Ø Philippines : Mindanao I. : Mt. Apo, 7°08N-125°09E, III-1985 (1 male, 1 female in coll. Mizunuma, Japan).

Ø Philippines : Mindanao I. : Surigao : Tandag, IV-1984 (2 males in coll. Mizunuma, Japan).

Ø Philippines : Mindanao I. : Bukidnon, 22-X-1991 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Mindanao I. : Baguio

Ø Philippines : Palawan, 9°37N-118°32E : Pinigisan

Ø Philippines : ? : Nueva Ecita Papaya

Ø Taiwan : Taichung Distr. (Lisan) (This locality is an error).

Ø Asia (TYPUS of Lucanus alces FABRICIUS, 1775 must be in Copenhagen Museum, Denmark).

Bibliography.

BOMANS H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des Coléoptères Lucanides). Lambillionea, CV(2):283-301.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis antilope VON ROTHENBURG, 1901[antilopa].

Von Rothenburg, 1901:25; Benesh, 1960:113; Maes, 1992:23.

Borneo*;

Sumatra.

Odontolabis brookeana (VOLLENHOVEN, 1861) [Lucanus, Calcodes] [brookeanus].

Vollenhoven, 1861:107, pl.6; Parry, 1864:15, pl.6; Leuthner, 1885:469, pl.95; Albers, 1894:167; Griffini, 1907:104-127; Heyne & Taschenberg, 1908:52, pl.7; Arrow, 1937:241, pl.1; Didier & Séguy, 1952:pl.45; Didier & Séguy, 1953:36; Benesh, 1960:113; Maes, 1980:210; Maes, 1981:5; Maes, 1981:145; Klausnitzer, 1982:43; Weng-Yew Tung, 1983:49, pl.7; Lacroix, 1984:98-99, pl.23; Kikuta, 1986:135; Nagai, 1986:3, 6, 9; Kawano, 1989:22; Téllez, 1990:40; Maes, 1992:23; Kawano, 1992:12; Mizunuma & Nagai, 1994:224, pl. 26.

Odontolabis brookeana ssp. brookeana (VOLLENHOVEN, 1861) [Lucanus] [brookeanus].

= Odontolabis ciliata ALBERS 1894 [India Or.].

? Malaya;

Borneo* : Sarawak, Sabah (Mt. Kinabalu, Mamut Copper Mine--Mt. Kinabalu), W. Kalimantan (Pontianak, Balaikarangan, Mt. Kenepai, Sambas, Mt. Bawang), ? (Labuan I.)

Borneo : Kinabalu, 1904 (1 male, 1 female in col. Ronald Cave, USA).

Borneo (2 males in col. Ronald Cave, USA).

Sumatra : N. Sumatra Prov. (Berastagi, Doloc Baros--Medan), Lampung Prov.,? (Langkat--Balei Gadjah, Deli).

Odontolabis brookeana ssp. coussementi MAES, 1981.

= Odontolabis brookeanus ssp. coussementi ab. brunnea MAES 1981 [Java].

Java*.

Odontolabis burmeisteri (HOPE, 1841) [Lucanus, Calcodes, Chalcodes].

= Anoplocnemus bicolor BURMEISTER 1847 [Mysore].

Hope, 1841:xxxiii; Hope, 1842:302; Hope, 1843:279, pl.13; Hope & Westwood, 1845:16; Burmeister, 1847:360; Parry, 1870:74; Leuthner, 1885:455, pl.88; Heyne & Taschenberg, 1908:52; Gravely, 1915:415; Arrow, 1950:193, pl.1, 16-17; Didier & Séguy, 1952:pl.46; Benesh, 1960:114; Lacroix, 1981:61; Sakaino, 1981:50; Lacroix, 1984:124-125, pl.34; Maes, 1992:23; Bomans, 1994:490; Mizunuma & Nagai, 1994:227, pl. 34.

India : Karnataka (Mysore*), Tamil Nadu (Nilgiri Hills, Anamalai Hills, Cinchona, Travancore, Coorg--Mercara, West Gaths near Trivandrum).

Note : The report of Endrodi (1968) of Nepal (Sun Kosi-Tal) is probably an error.

Odontolabis camela (OLIVIER, 1789).

Lucanus camelus OLIVIER, 1789:22-23, pl.5 [Loc. ?].

Olivier (ed. Illiger), 1800:74; Olivier (ed. Sturm), 1802:7, pl.4; Thunberg, 1806:185

Anoplocnemus alces FABRICIUS; Burmeister, 1847:359.

Odontolabis carinata LINNAEUS; Reiche, 1853:73.

Thomson, 1862:394; Parry, 1864:76

Odontolabis gouberti WATERHOUSE, 1876:172-173 [Mindoro].

Leuthner, 1885:456, pl.96; Nonfried, 1891:281; Séguy, 1954:87; Benesh, 1960:114; Lacroix, 1984:84-85, pl.17; Maes, 1992:24

Odontolabis camela (OLIVIER); Mizunuma & Nagai, 1994:225, pl. 27, 138, fig.158(1-8).

Odontolabis camela (OLIVIER); Bomans & Benoit, 2005:294.

Ø Philippines : S. Luzon I., IX/X-1989 (3 males in coll. Mizunuma, Japan).

Ø Philippines : S. Luzon I., 9-V-1992 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Luzon I., V-1975 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Luzon I., II-1984 (1 female in coll. Mizunuma, Japan).

Ø Philippines : Mindoro I., 12°58N-121°03E

Ø Philippines : Catanduanes I., 13°46N-124°16E, VIII-1993 (2 males in coll. Mizunuma, Japan).

Ø No Locality (TYPUS of Lucanus camelus OLIVIER, 1789 in MNHN, Paris).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis castelnaudi ssp. castelnaudi PARRY, 1862.

Odontolabis castelnaudi PARRY, 1862: 1862:108.

Odontolabis casternaudi.

Odontolabis castelnaui.

Parry, 1864:14, pl.1; Parry, 1870:74, pl.3; Leuthner, 1885:466, pl.95; Ritsema, 1913:179; Didier & Séguy, 1952:pl.47; Benesh, 1960:115; Mares & Lapacek, 1980; Lacroix, 1981:60; Maes, 1981:145; Klausnitzer, 1982:19; Weng-Yew Tung, 1983:49, pl.5; Lacroix, 1984:96-97; Nagai, 1986:2, 4-5, 7; Kawano, 1989:22; Maes,1992:24; Kawano, 1992:12; Kawano, 1993:14; Kawano, 1994:12

Odontolabis castelnaudi ssp. castelnaudi PARRY; Mizunuma & Nagai, 1994:13, fig. 21, p. 225, pl. 26-27.

Malaya : Perak (Taiping--4°51N-100°41E, Kampong Sahom), Pahang (Cameron Highlands);

Borneo : Sabah : Crocker Range, IV-2001, leg. Chew (1 male in col. Ronald Cave, USA).

Sumatra* : W. Sumatra Prov. (Padang), Lampung Prov.

Odontolabis castelnaudi ssp. inomatai MIZUNUMA, 1994.

Odontolabis castelnaudi ssp. inomatai MIZUNUMA in MIZUNUMA & NAGAI, 1994:10, 13, fig. 22, pl. 27, fig. 157(6-8) [Simeulue I.].

Sumatra : Aceh Prov. : Simeulue I., V-1992 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (female ALLOTYPUS in col. T. Mizunuma, Japan).

Sumatra : Aceh Prov. : Simeulue I., VI-1992 (3 males PARATYPUS in col. T. Mizunuma, Japan) (1 male PARATYPUS in col. H. Yasuoka, Japan).

Sumatra : Aceh Prov. : Simeulue I. = Simalur I. : Sibigo.

Odontolabis castelnaudi ssp. mentawaiensis MIZUNUMA, 1994.

Odontolabis castelnaudi ssp. mentawaiensis MIZUNUMA in MIZUNUMA & NAGAI, 1994:12-13, fig. 23, pl. 27, fig. 157(9-11) [North Pagai I.].

Sumatra : W. Sumatra Prov. : Mentawai Is. : Siberut I., VI-93 (2 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan)

Sumatra : W. Sumatra Prov. : Mentawai Is. : Sipora I., V-93(20 males PARATYPUS, 20 females PARATYPUS in col. T. Mizunuma, Japan) (2 males PARATYPUS, 2 females PARATYPUS in Kanagawa Prefectural Museum, Japan) (2 males PARATYPUS, 2 females PARATYPUS in col. H. Fujita, Japan) (4 males PARATYPUS, 4 females PARATYPUS in col. H. Yasuoka, Japan) (2 males PARATYPUS, 1 female PARATYPUS in col. T. Mizukami, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : North Pagai I., V-1992 (male HOLOTYPUS, female ALLOTYPUS in Kanagawa Prefectural Museum) (4 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : North Pagai I., VIII-1992 (10 males PARATYPUS, 2 females PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : South Pagai I., V-93 (40 males PARATYPUS in col. H. Fujita, Japan)

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis cuvera HOPE, 1842.

Saunders, 1839:177, pl.16; Hope, 1842:127; Hope, 1842:247; Hope, 1843:105, pl.10; Hope & Westwood, 1845:16; Burmeister, 1847:360; Westwood, 1848:54, pl.25-26; Thomson, 1862; Parry, 1864:14, 75, pl.1; Leuthner, 1885:450, 452-453, pl.91; Nonfried, 1891:281; Boileau, 1901:284; Meyer-Darcis, 1901:355; Mollenkamp, 1901:363; Boileau, 1902:204; Meyer-Darcis, 1902:410, pl.1; Mollenkamp, 1902:353; Mollenkamp, 1903:347; Zang, 1905:212; Mollenkamp, 1906:32; Heyne & Taschenberg, 1908:51, pl.7; Mollenkamp, 1909:9; Boileau, 1913:241; Pouillaude, 1913:334; Gravely, 1915:414; Vitalis de Salvaza, 1919:179; Kriesche, 1920:92; Kriesche, 1922:119; Arrow, 1950:189-191, 195, pl.16-18; Didier & Séguy, 1952:pl.48, 57; Didier & Séguy, 1953:60; Benesh, 1960:115-116, 119; Endrodi, 1968; Endrodi, 1971; Bomans, 1970:226; Numhauser, 1981:74; Lacroix, 1981:59-61; Lacroix, 1981:60-61; Sakaino, 1981:50; Klausnitzer, 1982:30, 42; Lacroix, 1984:126-141, 146-147, pl.35-39, 41; Bomans, 1988:299; Kawano, 1988:9; Bomans, 1989:6, 14; Kawano, 1989:22; Bomans, 1991:147; Maes, 1992:24, 27, Kawano, 1992:12; Kawano, 1992:12; Kawano, 1993:14; Ma, 1993:279; Bartolozzi & Sforzi, 1994:114; Mizunuma & Nagai, 1994:226-227, pl. 32-33; Kawano, 1997:454, 457, fig. 4.

Odontolabis cuvera ssp. cuvera HOPE, 1842 [Calcodes].

Calcodes cuvera HOPE, 1842.

= Lucanus bicolor SAUNDERS (nec OLIVIER) 1839 [Anoplocnemus].

= Odontolabis saundersi HOPE 1843 [saundersii] [Khasi Hills].

= Lucanus prinsepii HOPE & WESTWOOD 1845 [Assam].

= Lucanus delessertii HOPE & WESTWOOD (nec GUERIN-MENEVILLE) 1845.

= Odontolabis gazella THOMSON (nec FABRICIUS) 1862.

= Odontolabis burmeisteri ENDRODI (nec HOPE) 1968 & 1971.

Odontolabis cuvera HOPE; Kawano, 2003:503-511.

India : Sikkim

India : West Bengal (Darjeeling, Kurseong, Gopaldhara, Rungbong Valley, Lebong, Pashok, Mungphu)

India : Assam (Lamin)

India : Meghalaya : Khasi Hills : Shillong, 1975, ex. col. E. Coussement (2 males in col. Ronald Cave, USA).

India : Meghalaya : Cherrapunji

India : Meghalaya : Shillong

India : Manipur

India (2 females in col. Ronald Cave, USA).

Nepal

Bhutan : Padong

Bangladesh : Sylhet*.

Note : Gravely (1915) mention this species as rare in South India (Nilgiri Hills, Wynad), probably doesn't exist in south India.

Bibliography :

KAWANO K. (2003) Character displacement in Stag Beetles (Coleoptera: Lucanidae). Annals of the Entomological Society of America, 96(4):503-511.

Odontolabis cuvera ssp. rattii LACROIX, 1981.

Bhutan*.

Odontolabis cuvera ssp. mandibularis MOLLENKAMP, 1909.

India : Assam (Brahmaputra*).

Odontolabis cuvera ssp. lunulata LACROIX, 1984 [lunulatus].

India : Meghalaya (Shillong*).

Odontolabis cuvera ssp. gestroi BOILEAU, 1902.

= Odontolabis cuvera ssp. alticola MOLLENKAMP 1902 [Naga Hills].

India : Meghalaya (Shillong), Nagaland (Naga Hills, Kohima);

Myanmar = Burma : Kachin State (Kachin Kauri*), Rangoon State (Rangoon--16°48N-96°08E), Tenasserim State (Tavoy), ? (Bhamo, Nam Tamai Valley);

Thailand : Nakhon Ratchasima (Nakhon Ratchasima--15°01N-102°04E).

Odontolabis cuvera ssp. sinensis (WESTWOOD, 1848).

Lucanus sinensis WESTWOOD, 1848.

Odontolabis cuvera HOPE; Kawano, 2003:503-511.

Myanmar = Burma : S. Shan State (Loimwe)

China : Shanghai (Shanghai)

China : Sichuan : Mt. Simiang Shan

China : Sichuan : You Yang, 3-VI-1995, leg. L.J. Kim (2 males, 2 females in col. Ronald Cave, USA).

China : Hunan (Wuling Mts.)

China : N. Yunnan (Kut-Sin-Fou)

China : Guizhou = Kouy-Tcheou

China : Fujian (Mt. Wuyi Shan, Kuatun)

China : Hong Kong

China : Guangdong (Guangzhou = Canton)

China : (Lilong)

China : Hainan I. (Mt. Wuzhi Shan)

China*

Thailand = Siam : Chiang Mai (Chiang Mai--18°50N-98°57E)

Laos : Xieng-Khouang, P.K. Khouai (Vientiane), Ban Van Heua, Thonpheng, Ban Sam Thong.

Bibliography :

KAWANO K. (2003) Character displacement in Stag Beetles (Coleoptera: Lucanidae). Annals of the Entomological Society of America, 96(4):503-511.

Odontolabis cuvera ssp. boulouxi LACROIX, 1984.

China : Yunnan (Yunnan-Fou*).

Note : Probably the same as fallaciosus.

Odontolabis cuvera ssp. fallaciosa BOILEAU, 1901 [fallaciosus].

= Odontolabis fruhstorferi MEYER-DARCIS 1901 [Tonkin].

= Odontolabis salvazae POUILLAUDE 1913 [Tonkin].

China : Guangxi

Thailand : Lampang : Muang Pan

Laos : Samneua

Laos : 100 km N Vientiane : Vang Vieng, 5/26-V-2001, leg. L. Jingke (2 males in col. Ronald Cave, USA).

N. Vietnam : Annam

N. Vietnam : Tonkin : Sapa : Mt. Manson*

N. Vietnam : Tonkin : Ngan-son

N. Vietnam : Tonkin : Vietteri

N. Vietnam : Tonkin : Vinh Yen : Tam Dao

S. Vietnam : Dalat

Odontolabis mouhoti PARRY, 1864.

Odontolabis mouhoti ssp. mouhoti PARRY, 1864 [Calcodes] [mouhotii].

Calcodes mouhoti PARRY, 1864.

Odontolabis mouhoti PARRY; Kawano, 2003:503-511.

Myanmar = Burma : Karen State (Kawkareik--Dawna Range), Tenasserim State (Tavoy--Mintha Sakan);

Thailand = Siam* : Chiang Mai (Chiang Mai--18°50N-98°57E), Nakhon Ratchasima (Nakhon Ratchasima--15°01N-102°04E), Chantaburi (Khao Soi Dao);

Cambodia*;

Laos.

Note : For Mizunuma & Nagai (1994) O. mouhoti is a good species with O. elegans as a ssp. Possibly there are both good species, in some localities, there are both presents with O. fallaciosus, which can also be a good species.

Bibliography :

KAWANO K. (2003) Character displacement in Stag Beetles (Coleoptera: Lucanidae). Annals of the Entomological Society of America, 96(4):503-511.

Odontolabis mouhoti ssp. elegans MOLLENKAMP, 1901 [Calcodes].

Odontolabis cuvera HOPE; KAWANO, 1997.

Odontolabis mouhoti PARRY; Kawano, 2003:503-511.

Myanmar = Burma : Karen State (Karen Hills--Carin Cheba*), Mon Sate (Sukli);

Thailand : Chiang Rai (Wiangpapao), Chiang Mai (Chiang Mai--18°50N-98°57E, Fang), Tak (Umphang), Saraburi (Nakhon Nagok--Khao Yai National Park), Nakhon Ratchasima (Nakhon Ratchasima--15°01N-102°04E), Kanchanaburi (Khao Galak), ? (Nakhon Pathom--13°48N-100°03E);

? Vietnam;

Malaya.

Bibliography :

KAWANO K. (2003) Character displacement in Stag Beetles (Coleoptera: Lucanidae). Annals of the Entomological Society of America, 96(4):503-511.

Odontolabis dalmani ssp. dalmani (HOPE & WESTWOOD, 1845).

Lucanus dalmani HOPE & WESTWOOD, 1845:17.

Odontolabis dalmanni.

Odontolabis darmanni.

Odontolabis pubescens (BLANCHARD) PARRY, 1864:76 [Borneo].

Odontolabis gracilis (KAUP) DIDIER, 1932:41.

Thomson, 1862:394; Gemminger & Harold, 1868:948; Kaup, 1868:77; Parry, 1870:57; Leuthner, 1885:438-439, 442, pl.87, 88; Van De Poll, 1887:279, 280; Albers, 1889:234; Van De Poll, 1889:225; Nonfried, 1891:281; Mollenkamp, 1902:283; Mollenkamp, 1903:324; Heyne & Taschenberg, 1908:51, pl.7; Ritsema, 1908:51; Ritsema, 1913:179; Arrow, 1950:206, pl.19; Didier & Séguy, 1952:pl.49; Séguy, 1954:87; Benesh, 1960:115, 116, 117; Weinreich, 1971; De Lisle, 1976:276; Klausnitzer, 1982:42; Weng-Yew Tung, 1983:50, pl.5; Lacroix, 1984:65-67, 68-69, 70-71, 72-73, pl. 9, 10, 11, 12; Okajima, 1985; Nagai, 1986:2, 7, 8; Maes, 1992:24, 25; Ikeda, 1993; Kawano, 1993:12; Bomans, 1994:491, 495; Kawano, 1994:9.

Odontolabis dalmanni ssp. dalmanni (HOPE & WESTWOOD); Mizunuma & Nagai, 1994:15, fig. 27(a-e), 228-229, pl. 36-37, 138.

Araya, 1994:4; Kawano, 1997:454.

Myanmar = Burma : Tenasserim* State;

Thailand;

Malaya = Malacca : Perak (Taiping--4°51N-100°41E, Kampong Sahom), Pahang (Cameron Highlands);

Borneo : Anambas Is. : Tarempa I.

Borneo : Natuna Is. : Bunguran Besar I., 3°56N-108°15E

Borneo : Laut I., 4°43N-107°59E

Borneo : Sarawak

Borneo : Sabah : Crocker Range

Borneo : Sabah : Kundasang

Borneo : Sabah : Ranau : Mamut Copper Mines

Borneo : Sabah : Ranau, IV-2001, leg. Chew (1 male in col. Ronald Cave, USA).

Borneo : W. Kalimantan : Singkawang : Mt. Bawang

Borneo : W. Kalimantan : Sintang

Borneo : W. Kalimantan : Karimata I.

Borneo : Njabang

Sumatra : N. Sumatra Prov. (Medan), ? (Foerangie, Sirdang, Tambang Salida, Lobo Fakam, Ampugadang);

Java : Rookmaker.

Note : Boileau (1901) separated two forms without name from Sumatra and from Borneo.

Odontolabis dalmani ssp. simeuluensis MIZUNUMA, 1994.

Odontolabis gracilis KAUP, Ritsema, 1913.

Odontolabis dalmanni ssp. simeuluensis MIZUNUMA in MIZUNUMA & NAGAI, 1994:12-15, fig. 29(a-c, e), pl. 36, fig. 176 (10-11) [Simeulue I.].

Sumatra : Aceh Prov. : Simeulue I., III-1991 (male HOLOTYPUS, female ALLOTYPUS in Kanagawa Prefectural Museum, Japan) (3 males PARATYPUS, 3 female PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : Aceh Prov. : Simeulue I., VIII/X-1991 (1 male PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : Aceh Prov. : Simeulue I., V-1992 (8 males PARATYPUS in col. T. Mizunuma, Japan) (2 males PARATYPUS, 1 female PARATYPUS in col. H. Fujita, Japan).

Sumatra : Aceh Prov. : Simeulue I., III-1991 (3 males PARATYPUS, 2 females PARATYPUS in col. M. Baba, Japan).

Sumatra : Aceh Prov. : Simeulue I., VI-1992 (2 males PARATYPUS in col. M. Baba, Japan).

Sumatra : Aceh Prov. : Simeulue I. = Simalur I. (Sinabang).

Odontolabis dalmani ssp. gracilis KAUP, 1868.

Odontolabis gracilis KAUP, 1868 (Nias).

Odontolabis bellicosus CASTELNAU; Parry, 1870.

Odontolabis gracilis KAUP; Ritsema, 1891:238.

Odontolabis dalmanni ssp. gracilis KAUP; Mizunuma & Nagai, 1994:15, fig. 28(a, d-e).

Odontolabis gracilis KAUP; Bomans & Benoit, 2005:295.

Ø Sumatra : N. Sumatra Prov. : Nias I. (TYPUS of Odontolabis gracilis KAUP, 1868).

Ø Java (wrong locality)

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

RITSEMA C. (1891) Note XXXV: Two new species of the lucanoid genus Cyclommatus Parry. Notes Leyden Museum, XIII:235-238.

Odontolabis dalmani ssp. batuensis MIZUNUMA, 1994.

Odontolabis dalmanni ssp. batuensis MIZUNUMA in MIZUNUMA & NAGAI, 1994:14-15, fig. 33(a, c, e), pl. 36, fig. 176(12-15) [Simuku I.].

Sumatra : N. Sumatra Prov. : Batu Is. : Simuku I., III-1991, leg. S. Isogai (male HOLOTYPUS in col. T. Mizunuma, Japan).

Sumatra : N. Sumatra Prov. : Batu Is., I-1991 (female ALLOTYPUS, 1 male PARATYPUS in col. T. Mizunuma, Japan).

Odontolabis dalmani ssp. subita MIZUNUMA, 1994.

Odontolabis dalmanni ssp. subita MIZUNUMA in MIZUNUMA & NAGAI, 1994:14-15, fig. 35(a-b), pl. 36, fig. 176(16), pl. 138, fig. 176(17) [North Pagai I.].

Sumatra : W. Sumatra Prov. : Mentawai Is. : Siberut I., VI-1992 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : Siberut I., XI-1992 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : Siberut I., IV-1993 (2 males PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : N. Pagai I., II-1992 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : N. Pagai I., I-1993 (2 males PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : N. Pagai I., V-1993 (20 males PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : N. Pagai I., VIII-1993 (3 males PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawai Is. : S. Pagai I., III-1993 (25 males PARATYPUS in col. H. Fujita, Japan).

Odontolabis dalmani ssp. bellicosa (CASTELNAU, 1840).

Lucanus bellicosa CASTELNAU, 1840.

Odontolabis bellicosus.

Dorcus emarginatus DEJEAN, 1837 [nomen nudum] [Java].

Dorcus puncticeps (DE HAAN) DEJEAN, 1837 [nomen nudum] [Java].

Lucanus ursus CASTELNAU, 1840 [Java].

Lucanus serrifer HOPE & WESTWOOD, 1845 [Java].

Lucanus vishnu HOPE & WESTWOOD, 1845 [Java].

Anoplocnemus alces BURMEISTER (nec FABRICIUS) 1847.

Dejean, 1837:193; Castelnau, 1840:171, pl.16; Hope & Westwood, 1845:17; Burmeister, 1847:357; Reiche, 1853:72; Thomson, 1862:394; Parry, 1864:76; Gemminger & Harold, 1868:947; Leuthner, 1885:441, pl.88; Heyne & Taschenberg, 1908:51; Boileau, 1913:239; Oberthur & Houlbert, 1913:328; Didier & Séguy, 1952:pl.43-44; Benesh, 1960:113; Maes, 1981:1; Klausnitzer, 1982:14; Weng-Yew Tung, 1983:51, pl.6; Lacroix, 1984:74-75, pl.13; Maes, 1992:23; Mizunuma & Nagai, 1994:227, pl. 35.

Java* : Soekaboemi, Malang, Mt. Ardjoeno, Kawie Mts., Mt. Gede, Bali I.--8°20S--115°08E

Java (3 males, 2 females in col. Ronald Cave, USA).

[Not present in Sulawesi (=Celebes)]

Odontolabis dalmani ssp. celebensis LEUTHNER, 1885.

Odontolabis celebensis LEUTHNER, 1885 (Celebes).

Odontolabis dalmanni ssp. celebensis LEUTHNER; Mizunuma & Nagai, 1994:15, fig. 30(a-e).

Odontolabis celebensis LEUTHNER; Bomans & Benoit, 2005:295.

Sulawesi = Celebes* : Palu--0°46S-119°49E (Palolo), Palopo--2°55S-120°06E (Puncak), Mt. Bonthein, Todjamboe, Sangihe I. = Sanghir I.* (Tahuna), Banggai Is. (Peleng I.--1°18S-123°05E, Bangkulu I.), Wangi Wangi I.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis dalmani ssp. tahulandangensis MIZUNUMA, 1994.

Odontolabis dalmanni ssp. tahulandangensis MIZUNUMA in MIZUNUMA & NAGAI, 1994:14-15, fig. 31(a-c, e), pl. 37, fig. 176(33-36) [Tahulandang I.].

Sulawesi : Tahulandang I., VII-88, leg. Naomi Nishikawa (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (female ALLOTYPUS, 4 males PARATYPUS in col. T. Mizunuma, Japan).

Odontolabis dalmani ssp. sulaensis MIZUNUMA, 1994.

Odontolabis dalmanni ssp. sulaensis MIZUNUMA in MIZUNUMA & NAGAI, 1994:14-15, fig. 32(a-c, e), pl. 37, fig. 176(37-38) [Mangole I.].

Sulawesi = Celebes : Sula Is. : Mangole I., V-1992 (male HOLOTYPUS, female ALLOTYPUS in Kanagawa Prefectural Museum, Japan).

Sulawesi : Sula Is. : Mangole I., III-1992 (2 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).

Sulawesi = Celebes : Sula Is. : Taliabu I., VII-1988 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sulawesi = Celebes : Sula Is. : Taliabu I., III/V-1993 (10 males PARATYPUS, 7 females PARATYPUS in col. T. Mizunuma, Japan) (2 males PARATYPUS, 1 female PARATYPUS in col. H. Fujita, Japan).

Moluccas Is. : Halmahera I.--0°40N-128°E.

Odontolabis dalmani ssp. intermedia VAN DE POLL, 1889.

Odontolabis intermedius VAN DE POLL, 1889 (Palawan).

Odontolabis tarandus MOLLENKAMP, 1902 [Philippines].

Odontolabis femoralis ssp. palawanicus LACROIX, 1990 (Palawan).

Odontolabis dalmanni ssp. intermedia VAN DE POLL; Mizunuma & Nagai, 1994:15, fig. 34(a-b, d-e), p.228, pl.36, fig.176(18-21).

Odontolabis intermedia (VAN DE POLL); Bomans & Benoit, 2005:295.

Note : Bomans & Benoit (2005) consider that this taxon is not a subspecies of Odontolabis dalmani but a different species. Color is different and this toxon has not the pilosity on elytra that presents Odontolabis dalmani.

Ø Philippines : Sibuyan I.--12°25N-122°35E.

Ø Philippines : Romblon I.--12°26N-122°04E.

Ø Philippines : Panay I., 11°08N-122°23E : Alojian, I-1988 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Negros I. : Mt. Canlaon, 10°22N-123°07E, I-1987 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Negros I. : Mt. Canlaon, 10°22N-123°07E, XI-1989 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Cebu I., 10°18N-123°43E.

Ø Philippines : Mindanao I., 7°44N-125°02E.

Ø Philippines : Palawan I., 9°37N-118°32E (TYPUS of Odontolabis intermedius VAN DE POLL, 1889 probably in MNHN, Paris).

Ø Philippines : Palawan I., 9°37N-118°32E : S. Palawan, VIII-1979, ex. Coll. Cofais (TYPUS sell to Japan).

Ø Philippines : Palawan I., 9°37N-118°32E, IX-1989 (1 female in coll. Mizunuma, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis dalmani ssp. rudicae NAGAI & TSUKAMOTO, 2002.

Odontolabis dalmani ssp. rudicae NAGAI & TSUKAMOTO, 2002.

Odontolabis intermedia (VAN DE POLL); Bomans & Benoit, 2005:295.

Note : Make not a lot of sense to have a subspecies in all Philippines including Palawan and a subspecies in Mindoro. A good check to this situation is necessary. Also to check if intermedia is or not a subspecies of dalmani.

Ø Philippines : Mindoro I. : Oriental Province : Mt. Roosevelt, V-2000 (Male HOLOTYPUS in Entomological Laboratory in Ehime University, Ehime, Japan).

Bibliography.

Odontolabis delessertii (GUERIN-MENEVILLE, 1843) [Lucanus, Calcodes] [delesserti].

= Odontolabis cuvera PARRY (nec HOPE) 1863.

Guerin-Meneville, 1843:14, pl.12; Hope & Westwood, 1845:5, 31; Chenu, 1851:fig.2; Parry, 1863:447; Leuthner, 1885:454, pl.92; Heyne & Taschenberg, 1908:52; Arrow, 1950:192, pl.17; Didier & Séguy, 1952:pl.50; Didier & Séguy, 1953:36; Benesh, 1960:116; Sakaino, 1981:50; Klausnitzer, 1982:42; Lacroix, 1984:122-123, pl.33; Maes, 1992:24; Mizunuma & Nagai, 1994:227, pl. 34.

India : Kerala = Malabar (Wynad), Tamil Nadu : (Madurai--Shembaganur), Nilgiri Hills*, Travancore--High Range, Ponmudi, Palni Hills, Cuddapah).

Odontolabis eremicola MOLLENKAMP, 1905.

= Odontolabis sommeri var. cypri DIDIER & SEGUY 1953:37 [Borneo : Kinabalu].

= Odontolabis lowei ssp. aspericollis DE LISLE 1970 [Borneo].

Mollenkamp, 1905:7; Didier & Séguy, 1953:37; Benesh, 1960:117; De Lisle, 1970:98; Lacroix, 1984:104-105, pl.26; Nagai, 1986:6; Maes, 1992:25; Mizunuma & Nagai, 1994:224, 225, pl. 25.

Borneo : Sabah : Mt. Kinabalu*

Borneo : Sabah : Mt. Kinabalu : Headquarters

Borneo : Sabah : Crocker Range

Borneo : Sabah : Tawan, IV-2001, leg. Chew (1 male in col. Ronald Cave, USA).

? Sumatra : N. Sumatra Prov. : Medan

Odontolabis femoralis WATERHOUSE, 1887.

Waterhouse, 1887:446; Von Rothenburg, 1900:84; Mollenkamp, 1904:341; Boileau, 1913:240; Didier & Séguy, 1952:pl.52; Didier & Séguy, 1953:67; Benesh, 1960:117; Ratti & Lacroix, 1975:80-85; Weng-Yew Tung, 1983:50, pl.5; Lacroix, 1984:142-145, pl.40-41; Nagai, 1986:2, 6, 7, 12; Kawano, 1989:22; Lacroix, 1990:13-14, pl.1; Bomans, 1991:147; Maes, 1992:24-25; Kawano, 1992:12; Kawano, 1994:9; Mizunuma & Nagai, 1994:226, pl. 31; Kawano, 1997:454.

Odontolabis femoralis ssp. femoralis (WATERHOUSE, 1887) [Lucanus].

= Odontolabis chuai RATTI & LACROIX 1975 [Malaya : Cameron Highlands].

Thailand : ? (Chumphon);

Malaya : Pinang (Pinang I.--5°22N-100°12E), Perak* (Taiping--4°51N-100°41E, Kampong Sahom, Cameron Highlands--Taphh), Pahang (Cameron Highlands);

Sumatra.

Odontolabis femoralis ssp. waterstradti VON ROTHENBURG, 1900.

= Odontolabis waterstradti var. kinabaluensis MOLLENKAMP 1904 [Kinabalu].

Borneo : Sarawak : Mt. Murud

Borneo : Sabah : Mt. Kinabalu*

Borneo : Sabah : Mt. Kinabalu : Headquarters

Borneo : Sabah : Crocker Range

Borneo : Sabah : Ranau, IV-2001, leg. Chew (1 male in col. Ronald Cave, USA).

Borneo : Kalimantan : Mt. Poe

Odontolabis gazella (FABRICIUS, 1787).

Fabricius, 1787:1; Olivier, 1789:13-14, 22, pl.3-5; Herbst, 1790:313; Linnaeus (ed. Gmelin), 1790:1589; Fabricius, 1792:238; Olivier (ed. Illiger), 1800:61, 74; Fabricius, 1801:250; Olivier (ed. Sturm), 1802:3, 7, pl.2, 4; Thunberg, 1806:195, 204; Schonherr, 1817:322, 324; Burmeister, 1847:360; Thomson, 1862:395; Parry, 1864:77; Kaup, 1868:77; Parry, 1870:57; Leuthner, 1885:463, 465, pl.91, 96; Albers, 1886:28; Mollenkamp, 1904:347; Heyne & Taschenberg, 1908:52, pl.7; Kriesche, 1922:119; Didier, 1930:158; Didier & Séguy, 1953:36; Benesh, 1960:117; Weinreich, 1972; Maes, 1980:210; Maes, 1981:5-6; Klausnitzer, 1982:43; Weng-Yew Tung, 1983:51, pl.7; Lacroix, 1984:110-115, pl.29-31; Nagai, 1986:3, 8, 12; Bomans, 1991:147; Maes, 1992:25; Kawano, 1992:12; Mizunuma & Nagai, 1994:223-224, pl. 24; Kawano, 1997:454.

Odontolabis gazella ssp. gazella (FABRICIUS, 1787) [Lucanus].

= Lucanus bicolor OLIVIER 1789 [Loc. ?].

Thailand = Siam* : ? (Suratani);

Malaya = Malacca : Pinang (Pulo-Pinang), Perak (Taiping--4°51N-100°41E, Kampong Sahom), Pahang (Cameron Highlands);

Borneo : Natuna Is. (Bunguran Besar I.--3°56N-108°15E), Sarawak (Mt. Poi, Mt. Penrisson), Sabah (Mamut Copper Mine--Mt. Kinabalu), W. Kalimantan (Mt. Kenepai);

Sumatra : S. Sumatra Prov. (Mt. Dempo), ? (Tambang Salida); Karimata I.;

Philippines : Balabac I.--7°55N-116°58E.

Odontolabis gazella ssp. instabilis MOLLENKAMP, 1904.

Sumatra : N. Sumatra Prov. (Toba Lake*), ? (Pahang-Jalang).

Odontolabis gazella ssp. inaequalis KAUP, 1868.

Odontolabis inaequalis KAUP, 1868 (Nias).

Odontolabis inaequalis KAUP; Ritsema, 1891:238.

Odontolabis falcipedia DIDIER, 1930 [localité inconnue].

Odontolabis falcipedia DIDIER; Bomans & Benoit, 2005:295 (they note the dubious status of this species).

Ø Sumatra : N. Sumatra Prov. : Nias I. : Madjedja

Ø Sumatra : N. Sumatra Prov. : Nias I. : Goenoeng Sitoli

Ø Sumatra : N. Sumatra Prov. : Nias I. (TYPUS of Odontolabis inaequalis KAUP, 1868).

Ø Unknown Locality (male and female TYPUS of Odontolabis falcipedia DIDIER, 1930 probably in MNHN, Paris)

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

RITSEMA C. (1891) Note XXXV: Two new species of the lucanoid genus Cyclommatus Parry. Notes Leyden Museum, XIII:235-238.

Odontolabis gazella ssp. demoulini MAES, 1981.

Java*.

Odontolabis hitam NAGAI, 1986.

Nagai, 1986:3-5, 10-11; Mizunuma & Nagai, 1994:224, pl. 26.

N. Borneo : Sabah (Crocker Range* 16 mi NW of Keningau, Mamut Copper Mine* near Ranau--Mt. Kinabalu, Kundasan* near Ranau), Kalimantan (Mt. Poe);

Sumatra : W. Sumatra Prov. (Padang).

Note : Mizunuma & Nagai (1994) consider the populations of Kalimantan and Sumatra as distinct ssp.

Odontolabis imperialis MOLLENKAMP, 1904.

Odontolabis imperialis MOLLENKAMP, 1904:348 (Mt. Kinabalu).

Boileau, 1905:147

Odontolabis imperialis ssp. pallescens KRIESCHE, 1926:383.

Benesh, 1960:117; Lacroix, 1984:160-161, pl.47; Nagai, 1986:3, 8, 9; Maes, 1992:25; Kawano, 1992:12

Odontolabis imperialis MOLLENKAMP; Mizunuma & Nagai, 1994:226, pl. 31, fig.167(1-7).

Odontolabis imperialis MOLLENKAMP; Bomans & Benoit, 2005:295.

Ø Borneo : Sarawak : Mt. Serapi : near Kuching, 25-IV-1990 (1 male in coll. Mizunumai, Japan).

Ø Borneo : Sabah : Mt. Kinabalu (male TYPUS in MNHN, Paris).

Ø Borneo : Sabah : Crocker Range, 13-V-1987 (1 female in coll. Mizunumai, Japan).

Ø Borneo : Sabah : Crocker Range, 12-VI-1987 (1 male in coll. Mizunumai, Japan).

Ø Borneo : Sabah : Crocker Range, V-1989 (1 male in coll. Mizunumai, Japan).

Ø Borneo : Sabah : Crocker Range, IV-1990 (2 males in coll. Mizunumai, Japan).

Ø Borneo : Sabah : Crocker Range, V-1991 (1 male in coll. Mizunumai, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis imperialis ssp. komorii NAGAI & TSUKAMOTO, 2002.

Odontolabis imperialis ssp. komorii NAGAI & TSUKAMOTO, 2002 (Palawan).

Odontolabis imperialis MOLLENKAMP; Bomans & Benoit, 2005:295.

Ø Philippines : S. Palawan I. : Mt. Trus Maria Bataraza, V/VII-2002 (male HOLOTYPUS in Entomological Laboratory of Ehime University, Matsuyama, Japan).

Bibliography.

Odontolabis lacordairei (VOLLENHOVEN, 1861).

Lucanus lacordairei VOLLENHOVEN, 1861:104, pl.5 [Sumatra].

Odontolabis signatipennis MOLLENKAMP, 1906:170 [Sumatra].

Didier & Séguy, 1952:pl.53; Didier & Séguy, 1953:38; Benesh, 1960:118, 120; Weinreich, 1971; Weinreich, 1972; Klausnitzer, 1982:43; Lacroix, 1984:148-151, pl.42-43; Nagai, 1986:8; Kawano, 1989:22; Maes, 1992:25; Kawano, 1992:12.

Odontolabis lacordairei (VOLLENHOVEN); Mizunuma & Nagai, 1994:13, fig. 25(a, c-e), p. 225-226, pl. 29, 138.

Sumatra* : N. Sumatra Prov. (Berastagi, Dolok Merangir, ), S. Sumatra Prov. (Mt. Dempo), Lampung Prov., W. Sumatra Prov. (Padang, Mt. Singalan, Fort de Kock), ? (Sibolangit, Bukit Gompong);

?Borneo*.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis latipennis ssp. latipennis (HOPE & WESTWOOD, 1845).

Lucanus latipennis HOPE & WESTWOOD, 1845:17.

Anoplocnemus dejeani REICHE, 1852:23, pl.1 [Patria incognita].

Lucanus latipennis var. nigra VOLLENHOVEN, 1861:108.

Odontolabis styx BLANCHARD; Parry, 1864:76 [Singapur].

Gemminger & Harold, 1868:948; Leuthner, 1885:471-472, pl.96; Ritsema, 1913:177; Didier, 1930:147; Didier & Séguy, 1952:pl.54; Séguy, 1954:87; Benesh, 1960:117-119; Weinreich, 1972; Weng-Yew Tung, 1983:51, pl.6; Lacroix, 1984:116-121, pl.32; Nagai, 1986:3, 8, 12; Kawano, 1989:22; Lacroix, 1990:14; Maes, 1992:25-26; Kawano, 1992:12

Odontolabis latipennis (HOPE & WESTWOOD); Mizunuma & Nagai, 1994:223, pl. 24, fig.150(1-6).

Kawano, 1997:454.

Ø Malaya = Malacca : Kedah, 27-IV-1983 (1 female in coll. Mizunuma, Japan).

Ø Malaya = Malacca : Perak

Ø Malaya = Malacca : Pinang : Pinang I., 5°22N-100°12E = Prince of Wales I.

Ø Malaya = Malacca : Pahang : Cameron Highlands

Ø Malaya = Malacca : Johor

Ø Malaya = Malacca : Singapur

Ø Borneo : W. Kalimantan : Sintang

Ø Borneo : W. Kalimantan : Karimata I.

Ø Sumatra : Ache Prov. : Simeulue I. = Simalur I. : Sinabang

Ø Sumatra : Ache Prov. : Simeulue I. = Simalur I., 22-VIII-1992 (1 male in coll. Mizunuma, Japan).

Ø Sumatra : N. Sumatra Prov. : Berastagi, I-1983 (1 male in coll. Mizunuma, Japan).

Ø Sumatra : N. Sumatra Prov. : Medan

Ø Sumatra : Nias I.

Ø Sumatra : W. Sumatra Prov. : Padang, X-1991 (1 male in coll. Mizunuma, Japan).

Ø Sumatra : Siberut I.

Ø Sumatra : Bangka I. = Banka I., V-1991 (1 male in coll. Mizunuma, Japan).

Ø Sumatra : ? : Tambang Salida

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis latipennis ssp. planiceps DIDIER, 1930.

Odontolabis planiceps DIDIER, 1930:147.

Ø Moluccas Is. : Ambon I., 3°39S-128°07E = Amboina I. (TYPUS of Odontolabis planiceps DIDIER, 1930).

Odontolabis latipennis ssp. fratella LEUTHNER, 1885.

Odontolabis fratellus LEUTHNER, 1885:471-472, pl.96 (Luzon).

Odontolabis planiceps DIDIER; Benesh, 1958.

Odontolabis latipennis (HOPE & WESTWOOD); Mizunuma & Nagai, 1994:223, pl. 24, fig.150(1-6).

Odontolabis fratella LEUTHNER; Bomans & Benoit, 2005:295.

Ø Philippines : Luzon I. : Manilla, 14°35N-120°57E

Ø Philippines : Luzon I. (male and female TYPUS not localized).

Ø Philippines : Mindoro I.--12°58N-121°03E

Ø Philippines : Samar I., 11°49N-125°14E, IX-1991 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Leyte I., 10°54N-124°54E

Ø Philippines : Dinagat I., 10°09N-125°35E

Ø Philippines : Mindanao I. [Cotabato--7°14N-124°13E, Davao Prov. (Mt. McKinley)]

Ø Philippines : Palawan I., 9°37N-118°32E.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis leuthneri BOILEAU, 1897.

= Odontolabis leuthneri var. brunnea NAGEL 1926 [Sarawak : Mt. Murud].

Boileau, 1897:147; Von Rothenburg, 1900:92; Mollenkamp, 1904:341; Nagel, 1926:295, pl.12; Nagel, 1927:147; Didier & Séguy, 1952:pl.55; Benesh, 1960:118; De Lisle, 1976:276; Lacroix, 1984:pl.94, pl.21; Kikuta, 1986:135; Nagai, 1986:2, 6, 9; Maes, 1992:26; Kawano, 1994:9; Mizunuma & Nagai, 1994:223, pl. 24.

?Laos;

?Vietnam : Tonkin;

Borneo : Sarawak (Mt. Murud, Kalabit Country), Sabah (Mt. Kinabalu*, Headquarters--Mt. Kinabalu, Crocker Range).

Odontolabis lowei PARRY, 1873.

Parry, 1873:336, pl.3; Leuthner, 1885:470, pl.95; Van De Poll, 1890:159; Albers, 1894:165; Von Rothenburg, 1900:93; Mollenkamp, 1905:1; Zang, 1905:214; Griffini, 1906:111-139; Heyne & Taschenberg, 1908:52; Didier & Séguy, 1953:37; Benesh, 1960:118, 120; De Lisle, 1970; Maes, 1981:4-7; Maes, 1981:145-146; Klausnitzer, 1982:43; Lacroix, 1984:102-103, 108-109, pl.25, 28; Nagai, 1986:2, 5-6, 9, 11-12; Téllez, 1990:40; Maes, 1992:26; Kawano, 1992:12; Mizunuma & Nagai, 1994:224, pl.25.

Odontolabis lowei ssp. lowei PARRY, 1873.

= Odontolabis notarii LACROIX, 1984 [N. Borneo : Kinabalu, W. Sumatra].

Odontolabis lowei ssp. lowei f. chr. nubigena MOLLENKAMP, 1905 [Kinabalu].

Odontolabis lowei ssp. lowei f. chr. rufonotatus VON ROTHENBURG, 1900 [Borneo].

Borneo* : Sarawak (Mt. Murud), Brunei, Sabah (Mt. Kinabalu, Mamut Copper Mine--Mt. Kinabalu, Crocker Range), W. Kalimantan (Pontianak);

Sumatra : N. Sumatra Prov. (Doloc Baros State--Medan), W. Sumatra Prov., ? (Batak).

Odontolabis lowei ssp. juanitae MAES, 1995.

= Odontolabis cypri MAES (nec DIDIER & SEGUY) 1981 [Java].

Odontolabis lowei ssp. juanitae f. chr. nigra MAES 1981 [Java].

Java*.

Note : Nagai (1986) and Mizunuma & Nagai (1994) consider O. lowei as a ssp. of O. sommeri but we consider that's not possible because both are present in Borneo and Sumatra.

Odontolabis ludekingi (VOLLENHOVEN, 1861) [Lucanus].

Vollenhoven, 1861:105, pl.5; Parry, 1864:13, pl.2; Leuthner, 1885:460, pl.93-94; Mollenkamp, 1906:170; Van Roon, 1906:271, 276, pl.13; Didier & Séguy, 1952:pl.56; Benesh, 1960:118; Klausnitzer, 1982:43; Lacroix, 1984:152-155, pl.44-45; Bomans, 1990:176; Maes, 1992:26; Kawano, 1992:12; Mizunuma & Nagai, 1994:226, pl.30; Kawano, 1997:454.

Odontolabis ludekingi ssp. ludekingi (VOLLENHOVEN, 1861) [Lucanus].

Malaya = Malacca;

Sumatra* : N. Sumatra Prov. (Dolok Merangir), W. Sumatra Prov. (Padang, Padangpandjang, Fort de Kock, Mt. Singalan), Jambi Prov., Bengkulu Prov. (Tambang Sawah), Lampung Prov., ? (Ajer Mantcior, Bandong-Sijthoff).

Odontolabis ludekingi ssp. monticola MOLLENKAMP, 1906.

= Odontolabis ludekingi var. robustus VAN ROON 1906 [robusta] [Palembang].

Sumatra* : W. Sumatra Prov., S. Sumatra Prov. (Palembang), ? (Widdagedo).

Bibliography.

ARAYA K. (2000) An account of a visit to European Museums. 2. The lucanid specimens in the van Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke Historie, Leiden. Gekkan-Mushi, 350:4-16.

Odontolabis macrocephala LACROIX, 1984 [macrocephalus].

Lacroix, 1984:80-81, pl.15; Bomans, 1991:147; Maes, 1992:26; Mizunuma & Nagai, 1994:229, pl. 38.

China : Guangxi = Kouangsi*, Guangdong;

Thailand : Chiang Rai (Wiangpapao), Chiang Mai (Chiang Mai--18°50N-98°57E);

Vietnam : Tonkin (Hanoï*, Tam-Dao*).

Odontolabis micros DE LISLE, 1970.

De Lisle, 1970:96; Lacroix, 1981:59; Lacroix, 1984:90-91, pl.19; Maes, 1992:26; Kawano, 1994:12; Mizunuma & Nagai, 1994:224, pl.26.

C. & S. Sulawesi = Celebes : Todjamboe*, Palopo--2°55S-120°06E (Puncak).

Odontolabis mollenkampi FRUHSTORFER, 1898.

Fruhstorfer, 1898:167; Benesh, 1960:119; Lacroix, 1984:156-157, pl.46; Maes, 1992:26.

C. Sumatra*.

Odontolabis pareoxa BOMANS & RATTI, 1973 [pareoxus].

Bomans & Ratti, 1973:222; Lacroix, 1984:82-83, pl.16; Maes, 1992:26.

India : Nagaland (Naga Hills*).

Odontolabis picea BOMANS, 1986 [piceus].

Bomans, 1986:385; Maes, 1992:26.

Sumatra : S. Sumatra Prov. (Palembang).

Odontolabis platynota (HOPE & WESTWOOD, 1845).

Hope & Westwood, 1845:18; Saunders, 1854:49, pl.3; Westwood, 1855:201, pl.10; Thomson, 1862:394; Parry, 1864:77; Leuthner, 1885:435, pl.88; Heyne & Taschenberg, 1908:51, pl.7; Didier, 1927:271; Arrow, 1950:201, pl.21; Benesh, 1960:119; Bomans, 1970:226; Klausnitzer, 1982:45; Lacroix, 1984:78-79, pl.15; Bomans, 1989:6; Maes, 1992:26; Ma, 1993:279; Mizunuma & Nagai, 1994:229, pl. 38.; Kawano, 1997:454

Odontolabis platynota ssp. platynota (HOPE & WESTWOOD, 1845) [Lucanus, Calcodes] [platynotus, platinota] [East India].

= Odontolabris emarginata (REICHE) SAUNDERS 1854 [emarginatus] [China].

= Odontolabris evansii WESTWOOD 1855 [China].

? India : Assam;

N. Myanmar = Burma : Karen State (Karen Hills--Cheba);

China : Shanghai (Shanghai), Sichuan (Miyi), Hunan (Wuling Mts.), Yunnan, Guizhou, Zhejiang, Fujian (Kuatun), Hong Kong;

Laos : Xieng-Khouang, Parkading, Ban Van Heua, P.K. Khouei, Tonpheng.

Odontolabis platynota ssp. coomani DIDIER, 1927.

N. Vietnam : Tonkin (Hoa Binh*, Tam Dao near Vinh Yen).

Odontolabis quadrimaculata KRIESCHE, 1920 [quadrimaculatus].

Kriesche, 1920:92; Benesh, 1960:119; Maes, 1992:26.

Sumatra : (Sarik Alahan Pandschang*).

Odontolabis relucens MOLLENKAMP, 1900.

Mollenkamp, 1900:45; Mollenkamp, 1903:348; Benesh, 1960:119; Lacroix, 1984:162-163, pl.47; Maes, 1992:27.

Note : This species could have been described on an immature of O. imperialis (Lacroix, 1984).

Sumatra*.

Odontolabis siva (HOPE & WESTWOOD, 1845).

Hope & Westwood, 1845:5, 16; Reiche, 1853:72; Thomson, 1862:394; Parry, 1864:76; Leuthner, 1885:436, pl.86; Nonfried, 1891:281; Boileau, 1905:47; Heyne & Taschenberg, 1908:51, pl.7; Mollenkamp, 1909:9; Boileau, 1913:238; Gravely, 1915:415; Kriesche, 1922:119; Miwa, 1932:90; Miwa, 1936:2; Arrow, 1937:241, pl.1; Arrow, 1943:134; Arrow, 1950:200, pl.19; Arrow, 1951:pl.15; Didier & Séguy, 1952;pl.58-59; Didier & Séguy, 1953:37, 101; Séguy, 1954:87; Benesh, 1960:119; Bomans, 1970:226; Klausnitzer, 1982:29; Lacroix, 1984:76-77, pl.14; Biswas & Chatterjee, 1985:182; Bomans, 1989:6; Kojima, 1990:4-11; Bomans, 1991:148; Maes, 1992:27; Kawano, 1993:12; Ma, 1993:279; Sakaino & Ochi, 1993:20; Sakaino, Yu & Chu, 1994:123; Mizunuma & Nagai, 1994:227, pl. 34; Kawano, 1997:454.

Odontolabis siva ssp. siva (HOPE & WESTWOOD, 1845) [Lucanus, Calcodes].

= Odontolabis bellicosus REICHE (nec CASTELNAU) 1853.

= Odontolabis carinatus PARRY (nec LINNAEUS) 1864.

= Neolucanus alces var. portax DIDIER & SEGUY 1952 [Inde].

= Odontolabis parryi ab. mingianus DIDIER & SEGUY 1953 [Sikkim].

India : Sikkim, West Bengal (Darjeeling, Rungeer Valley, Mangpu, Gopaldhara, Rungbong Valley, Kurseong, Duars-Buxa), Arunachal Pradesh (Namdapha, Miao), Assam (Sibsagar), Meghalaya (Shillong, Khasi Hills*), Nagaland (Patkai Hills); Bangladesh : Sylhet*, Maini Mukh, Chittagong Hill;

Nepal;

Bhutan : Pedong;

Myanmar = Burma : Shan State (Muong Yoo);

China : Hunan (Wuling Mts.), Yunnan, Fujian (Kuatun), Hainan I.;

Thailand : Chiang Rai (Mae Jan, Wiangpapao), Chiang Mai (Mt. Doi Pui, Samoeng, Mt. Doi Saket), Tak (Umphang), Mukdaharn (Phu Pahn), Nakhon Nayok (Wang ta Krai), Petchaburi (Lomgao);

Laos : P.K.Khouai, Vientiane, Parkading, Pakse, Houei Kong, Muong Soui, Ban Van Heua, Thonpheng, Sayabouri;

Vietnam : Tonkin (Tam Dao near Vinh Yen), Annam;

Malaya = Malacca.

Odontolabis siva ssp. parryi BOILEAU, 1905.

= Odontolabis carinatus MIWA (nec LINNAEUS) 1927.

= Calcodes chinensis ARROW 1943 (new name for parryi).

Taiwan* : Taipei Distr. (Mt. Tatun Shan), Hualien Distr. (Taihorin), Nantou Distr. (Sunkang, Puli = Hori), ? (Hoozan, Kosempo, Suisharyo, Sokutsu, Kankau, Jisugetsutan, Shikikun, Urai, Thaiheizan, Taipin, Taihoku, Sakahen in Nôkô, Kobayashi to Ratô, Baibara), Lanyu I.--22°00N-121°32E, Lutao I.--22°39N-121°28E.

Odontolabis somneri PARRY, 1862.

Parry, 1862:108; Parry, 1864:16, pl.6; Gestro, 1881:313; Leuthner, 1885:467, pl.95; Van De Poll, 1890:160; Von Rothenburg, 1900:1, 59, 61; Zang, 1905:213; Didier & Séguy, 1952:pl.60; Benesh, 1960:120; Bomans, 1970:314; Weinreich, 1971; De Lisle, 1976:276; Maes, 1980:210; Maes, 1981:3; Maes, 1981:145; Klausnitzer, 1982:28; Weng-Yew Tung, 1983:51, pl.6; Lacroix, 1984:80, 100-101, 106-107, pl.24, 27; Bomans, 1986:385; Nagai, 1986:5-6; Kawano, 1989:21; Kawano, 1989:22; Maes, 1992:27; Kawano, 1992:12; Bomans, 1994:186; Mizunuma & Nagai, 1994:224, pl. 25-26; Kawano, 1997:454.

Odontolabis somneri ssp. pahangensis BOMANS, 1970.

Malaya : Perak (Kampong Sahom), Pahang (Cameron Highlands*, Fraser's Hills).

Odontolabis somneri ssp. somneri PARRY, 1862 [sommeri] [Philippines : Manilla, error].

= Odontolabis lowei GESTRO (nec PARRY) 1881.

Borneo : Sabah (Mt. Kinabalu, Kinabalu National Park), Kalimantan;

Sumatra : N. Sumatra Prov. (Bander Baru, Berastagi, Medan--Dolok Baros Estate, Dolok Merangir), W. Sumatra Prov. (Solok), S. Sumatra Prov. (Benkoelen), Bangka I. = Banka I., Lampung Prov., ? (Langkat--Balei Gadjah, Deli, Tambang Salida).

Odontolabis spectabilis BOILEAU, 1902.

Odontolabis lacordairei LEUTHNER (nec VOLLENHOVEN) 1885:461, pl. 94.

Odontolabis spectabilis BOILEAU, 1902:38-39, pl.1 [Sumatra].

Odontolabis spectabilis BOILEAU, 1902:204 [Sumatra].

Mollenkamp, 1906:170; Didier & Séguy, 1952:pl.53; Didier & Séguy, 1953:38; Benesh, 1960:118, 120; Weinreich, 1971; Weinreich, 1972; Klausnitzer, 1982:43; Lacroix, 1984:148-151, pl.42-43; Nagai, 1986:8; Kawano, 1989:22; Maes, 1992:25; Kawano, 1992:12.

Odontolabis spectabilis BOILEAU; Mizunuma & Nagai, 1994:13, fig. 26(a-e), p. 225-226, pl. 29, 138.

Sumatra* : N. Sumatra Prov. (Berastagi), W. Sumatra Prov. (Padang), ? (Sitahoan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Odontolabis stevensi THOMSON, 1862.

Thomson, 1862:414; Parry, 1864:76, pl.5; Deyrolle, 1865:26, pl.1; Parry, 1870:56; Parry, 1872:84, pl.1; Oberthur, 1879:566; Leuthner, 1885:447-449, pl.90; Schaufuss, 1887:117; Heyne & Taschenberg, 1908:51; Kriesche, 1922:119; Didier & Séguy, 1952:pl.51, 61; Didier & Séguy, 1953:61; Benesh, 1960:116, 120; Lacroix, 1981:59-60; Klausnitzer, 1982:42; Lacroix, 1984:86-89, 92-93, pl.18-19, 20; Maes, 1992:25, 27; Bomans, 1994:491; Mizunuma & Nagai, 1994:225, pl. 28.

Odontolabis stevensi ssp. stevensi THOMSON, 1862 [stevensii].

= Odontolabis stevensi var. bidentata LEUTHNER 1885.

N. & C. Sulawesi = Celebes : Manado*--1°31N-124°47E, Tondano--1°14N-124°51E, Palu--0°46S-119°49E (Palolo), Todjamboe.

Odontolabis stevensi ssp. limbata KRIESCHE, 1922 [limbatus].

C. & S. Sulawesi = Celebes* : Palopo--2°55S-120°06E (Puncak).

Odontolabis stevensi ssp. duivenbodei DEYROLLE, 1865 [Celebes : Manado, error].

Sulawesi = Celebes : Sangihe I. = Sanghir I. (Tahuna);

Lesser Sunda Is. : Timor I.--9°13S-124°56E.

Odontolabis versicolor (DIDIER, 1931) [Neolucanus, Calcodes, Chalcodes].

Didier, 1931:228; Arrow, 1935:108; Arrow, 1950:188, pl.20; Didier & Séguy, 1953:94; Benesh, 1960:120; Lacroix, 1984:51-53; Bartolozzi & Bomans, 1990:433; Maes, 1992:27; Mizunuma & Nagai, 1994:229, pl. 38, 139.

India : Karnataka (Mysore, Sagar), Tamil Nadu (Nilgiri Hills, Kanara*, Mundakayam, Travancore, Tinnevely, Madras).

Odontolabis vollenhoveni PARRY, 1864 [vollenhovii].

Parry, 1864:13, pl.8; Leuthner, 1885:459, pl.93; Heyne & Taschenberg, 1908:52; Didier & Séguy, 1952:pl.62; Didier & Séguy, 1953:62; Benesh, 1960:121; Klausnitzer, 1982:43; Lacroix, 1984:158-159, pl.46; Nagai, 1986:3, 6, 9; Maes, 1992:27; Kawano, 1992:12; Mizunuma & Nagai, 1994:226, pl. 30.

Borneo : Sabah : Kinabalu

Borneo : Sabah : Crocker Range

Borneo : Sabah : Trus Madi, III-2001, leg. Chew (1 male in col. Ronald Cave, USA).

Borneo*

Odontolabis wollastoni PARRY, 1864 [wallastoni].

= Odontolabis wollastoni var. pectinata LEUTHNER 1885.

Parry, 1864:14, pl.2; Leuthner, 1885:457, pl.93; Heyne & Taschenberg, 1908:52, pl.7; Didier & Séguy, 1952:pl.63; Didier & Séguy, 1953:63; Benesh, 1960:121; Weinreich, 1972; Klausnitzer, 1982:43; Weng-Yew Tung, 1983:52, pl.6; Lacroix, 1984:164, pl.48; Maes, 1992:27; Kawano, 1992:12; Haitlinger, 1994:196; Mizunuma & Nagai, 1994:226, pl. 30; Suzuki, 1995:17; Kawano, 1997:454.

Malaya = Malacca* : Cameron Highlands;

Sumatra : N. Sumatra Prov. (Berastagi, Medan--Doloc Baros Estate), W. Sumatra Prov., S. Sumatra Prov. (Mt. Dempo, Palembang, Lahat), ? (Sibolangit).

Natural enemies : Acarina : Canestriniidae : Sandrophela hugoni.

Odontolabis yasuokai MIZUNUMA, 1994.

Odontolabis yasuokai MIZUNUMA in MIZUNUMA & NAGAI, 1994:12-13, fig. 24(a, c-e), p. 226, pl. 29, fig. 162(1-5) [Mt. Dempo].

Sumatra : S. Sumatra Prov. : Mt. Dempo, VI-1988 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, III-1986 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, VII-1986 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, XII-1987 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, XII-1988 (2 males PARATYPUS in col. M. Baba, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, II-1991 (1 male PARATYPUS, 2 females PARATYPUS in col. M. Baba, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, IV-V-1991 (20 males PARATYPUS, 5 females PARATYPUS in col. T. Mizunuma, Japan) (7 males PARATYPUS, 2 females PARATYPUS in col. H. Yasuoka, Japan) (2 males PARATYPUS in col. T. Mizukami, Japan) (2 males PARATYPUS in col. H. Fujita, Japan).

Sumatra : S. Sumatra Prov. : Mt. Dempo, XII-1991 (2 males PARATYPUS in col. M. Baba, Japan).

Sumatra : S. Sumatra Prov. : Lampung Prov., III-86 (female ALLOTYPUS in col. T. Mizunuma, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

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