Neolucanus THOMSON, 1862

(G. = Odontolabis baladeva)

= Anodontolabis·PARRY 1863 (G. = Odontolabis baladeva)


Neolucanus armatus LACROIX, 1972.

Lacroix, 1972:111; Mizunuma, 1985:15-16; Maes, 1992:27.

Note : Pierre Benoit (in litteris) consider this as a synonym of Neolucanus delicatus Didier.

Vietnam : Tonkin (Tam Dao*--NE Hanoï).

Neolucanus aterrimus WEINREICH, 1959.

= Neolucanus interrimus LACROIX (nec WEINREICH) 1988.

Weinreich, 1959; Lacroix, 1988:6; Maes, 1992:27.

China : Xizang = Thibet (Giufu-shan*).

Neolucanus baladeva (HOPE, 1842) [Odontolabis, Calcodes] [baladera, baladevus].

= Lucanus angulatus HOPE & WESTWOOD 1845 [Neolucanus] [Assam].

= Neolucanus lama LEUTHNER (nec OLIVIER) 1885.

= Neolucanus waterhousei BOILEAU 1899 [Sikkim].

= Neolucanus celebensis MOLLENKAMP 1900 [Celebes, error].

= Neolucanus ollenbachi DIDIER 1930 [Naga Hills].

= Calcodes celebicus ARROW 1943 (new name for celebensis).

Note : Mizunuma & Nagai (1994) consider angulatus as a good species.

Hope, 1842:127; Hope, 1842:247; Hope, 1843:105; Hope & Westwood, 1845:17; Parry, 1870:58; Leuthner, 1885:430, pl.85; Boileau, 1899:178; Mollenkamp, 1900:46; Mollenkamp, 1903:349; Heyne & Taschenberg, 1908:52, pl.7-8; Boileau, 1913:247; Pouillaude, 1914:40; Gravely, 1915:414; Didier, 1930:131; Nagel, 1930:90; Miwa, 1932:96; Arrow, 1937:243; Arrow, 1943:34, 135; Arrow, 1950:204, pl.18; Didier & Séguy, 1953:34-35; Benesh, 1960:122; Sakaino, 1981:50; Klausnitzer, 1982:45; Lacroix, 1988:6; Maes, 1992:28; Bartolozzi & Sforzi, 1994:114; Mizunuma & Nagai, 1994:222-223, pl. 23; Kawano, 1997:454.

India : Sikkim, West Bengal (Darjeeling, Mangpu, Kurseong, Lebong, Debrepani), Assam, Meghalaya (Jaintai Hills, Shillong, Khasi Hills), Nagaland (Naga Hills), Manipur;

Nepal;

Bhutan : Pedong;

Bangladesh : Sylhet*;

Myanmar = Burma : Karen State (Cheba--Karen Hills);

China : Yunnan;

Thailand;

Taiwan : ? (Kosempo, Chip Chip).

Note : For Mizunuma & Nagai (1994:223) N. angulatus is a valid species from NE India (Darjeeling).

Bibliography.

ARAYA K. (2000) An account of a visit to European Museums. 2. The lucanid specimens in the van Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke Historie, Leiden. Gekkan-Mushi, 350:4-16.

 

Neolucanus borneensis HOULBERT, 1914 [Odontolabis].

= Neolucanus borneensis var. niger NAGEL 1926 [Mt. Murud].

Houlbert, 1914:279; Nagel, 1926:249, 294; Nagel, 1927:146; Benesh, 1960:122; Nagai, 1986:10; Maes, 1992:28.

Borneo : Sarawak (Mt. Murud), W. Kalimantan (Pontianak*, Sintang River*).

Neolucanus brevis BOILEAU, 1899.

Boileau, 1899:197; Mollenkamp, 1900:46; Zang, 1905:212; Arrow, 1950:203, pl.21; Benesh, 1960:123; Bomans, 1970:226; Lacroix, 1972:119; Téllez, 1990:39; Bomans, 1991:146; Maes, 1992:28, 30; Bomans, 1994:490; Mizunuma & Nagai, 1994:220, pl. 21; Kawano, 1997:454; Kawano, 1997:25.

Neolucanus brevis ssp. brevis BOILEAU, 1899 [Calcodes] [brebis, breves].

= Neolucanus birmaensis MOLLENKAMP 1900.

Myanmar = Burma : Kachin State (Kauri--Kachin Hills*), ? (Thandaung);

China : Yunnan;

Thailand : Chiang Rai (Wiangpapao), Chiang Mai (Chiang Mai--18°50N-98°57E, Mt. Doi Inthanon--18°26N-98°17E), Loei (Dansai), Petchaburi (Khao Khor);

Vietnam: Tonkin.

Neolucanus brevis ssp. rondoni LACROIX, 1972.

Thailand : Chiang Mai (Nan);

Laos : Tonpheng*, Ban Van Heua, Phou Khao Khouei (Ritaville* 40 km NE Vientiane), Phou Soai Dao.

 

Neolucanus brochieri BOMANS & MIYASHITA, 1997.

Neolucanus brochieri BOMANS & MIYASHITA, 1997:3, figs. 2, 3, pl. fogs. 3-4 [Kachin Prov.].

Myanmar :

Kachin Prov., VIII-96, leg. P. Sukkit (male HOLOTYPUS of Neolucanus brochieri BOMAN & MIYASHITA, female ALLOTYPUS in coll. Miyashita, Japan) (3 males PARATYPUS in coll. Miyashita, Japan; Bomans, Natural History Museum, London; Arnaud, Francia)

Neolucanus brochieri (adapted from Bomans & Miyashita).

 

Bibliography :

BOMAN H. & MIYASHITA T. (1997) Description de trois nouvelles espèces de Lucanidae de nord Birmanie. 91 ème contribution à l'étude des Coléoptères Lucanides. Besoiro, 4:2-3.

 

Neolucanus castanopterus (HOPE, 1831).

Hope, 1831:22; Hope & Westwood, 1845:18; Burmeister, 1847:360; Westwood, 1848:22, pl.10, pl.26; Leuthner, 1885:423, pl.84; Mollenkamp, 1907:109; Heyne & Taschenberg, 1908:52, pl.7; Mollenkamp, 1909:134; Mollenkamp, 1913:128; Boileau, 1914:73, 105; Gravely, 1915:414; Kriesche, 1922:123; Didier, 1930:146; Miwa, 1934:328; Nagel, 1941:54; Arrow, 1943:135; Arrow, 1950:197, pl.21; Didier & Séguy, 1953:60; Benesh, 1960:123, 124; Endrodi, 1971:10; De Lisle, 1976:273-274; Klausnitzer, 1982:43; Lacroix, 1988:6; Maes, 1992:28; Ma, 1993:279; Bartolozzi & Sforzi, 1994:114; Mizunuma & Nagai, 1994:219, pl. 20.

Neolucanus castanopterus ssp. castanopterus (HOPE, 1831) [Lucanus, Calcodes].

= Anoplocnemus bicolor BURMEISTER 1847 [Nepal].

= Neolucanus elongatus MOLLENKAMP 1907 (Perhaps synonym of N. swinhoei?) (perhaps a ssp. from Taiwan?).

= Neolucanus cephalotes MOLLENKAMP 1909 [Taiwan] (Perhaps ssp. from Taiwan?).

= Neolucanus bifoveolatus MOLLENKAMP 1913 [Taiwan] (Perhaps ssp. from Taiwan?).

= Neolucanus melas DIDIER 1930 [Manipur] (seems to be a dark ssp. of Manipur).

= Neolucanus parvus NAGEL 1941 [Shillong].

India : Uttar Pradesh (Almora, Ramnee), Sikkim (Shamdang, Gangtok), West Bengal (Darjeeling, Kurseong, Mungphu, Singla, Duars-Buxa), Assam (Sibsagar), Meghalaya (Cherrapunji, Shillong, Khasi Hills), Nagaland, Manipur;

Nepal* : Katmandu, Sun Kosi-Tal, Jiri, Jubing;

Bhutan : Maria Basti, Chukka;

? Taiwan.

Neolucanus castanopterus ssp. elongatulus MOLLENKAMP, 1907.

= Neolucanus pallidus BOILEAU 1914 [Yunnan-Sen].

China : Hunan (Wuling Mts.), Yunnan* (Sse-Tsong, Kouang-Si-Hien, Yunnan-Sen, Yunnan-Fou).

Neolucanus castanopterus ssp. flavipennis BOILEAU 1914.

Myanmar = Burma = Birmanie* : ? (Ruby Mines);

Thailand : Chiang Mai (Mt. Doi Inthanon--18°26N-98°17E);

Vietnam : Tonkin.

Neolucanus cingulatus PARRY, 1864.

Parry, 1864:20, pl.4; Gestro, 1881:314; Leuthner, 1885:421, 422, 428, pl.84, 85; Mollenkamp, 1897:145; Mollenkamp, 1905:7; Benesh, 1960:123, 124; Bomans, 1973:279; Nagai, 1986:3, 10; Maes, 1992:28, 29; Mizunuma & Nagai, 1994:220, pl. 21; Kawano, 1997:454; Kawano, 1997:25.

Neolucanus cingulatus ssp. cingulatus PARRY, 1864.

Malaya = Malacca* : Perak (Taiping--4°51N-100°41E), Pahang (Cameron Highlands, Pantai Remis).

Neolucanus cingulatus ssp. lansbergei LEUTHNER, 1885.

Sumatra* : W. Sumatra Prov. (Fort de Koch-Padang, Bukittinggi), ? (Harau Valley).

Neolucanus cingulatus ssp. muntjac GESTRO, 1881 [Odontolabis].

Borneo : Natuna Is. (Bunguran Besar I.--3°56N-108°15E), Sarawak* (Mt. Serapi near Kuching), W. Kalimantan (Sanggau).

Neolucanus curvidens LACROIX, 1978.

Lacroix, 1978:267; Maes, 1992:28.

China : Guizhou = Kouy Tcheou*.

Note : Perhaps a ssp. of N. sinicus.

Neolucanus delicatus DIDIER, 1927.

Didier, 1927:144; Didier & Séguy, 1953:94; Benesh, 1960:123; Mizunuma, 1985:15-16; Maes, 1992:28; Kawano, 1994:9; Mizunuma & Nagai, 1994:217, 311, pl.18, 149; Baba:1995:7-11; Kawano, 1997:454; Kawano, 1997:25.

Vietnam : Annam (Col des Nuages* near of Tourane), Tonkin (Tam Dao near Vinh Yen).

Neolucanus didieri LACROIX, 1972.

Lacroix, 1972:117; Maes, 1992:28.

Vietnam : Annam (Djiring*).

Neolucanus diffusus BOMANS, 1989.

Bomans, 1989:5, 12-14.

China : Fujian (Kuatun*).

Neolucanus donckieri DIDIER, 1926.

Didier, 1926:211; Didier, 1929:85; Benesh, 1960:123; Lacroix, 1988:6; Maes, 1992:28; Mizunuma & Nagai, 1994:311, pl.149.

China : Yunnan (Yunnan-Sen*);

Laos*.

 

Neolucanus doro MIZUNUMAI, 1994.

Neolucanus doro MIZUNUMAI in MIZUNUMA & NAGAI, 1994:8, 11, fig. 12(a-d), 219, pl. 20, fig. 129(1-8).

Araya, 1996:21.

Taiwan : Nantou Distr. : Mt. Hehuanmou, 2/21-X-86 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (20 males PARATYPUS, 10 females PARATYPUS in col. T. Mizunuma, Japan).

Taiwan : Nantou Distr. : Mt. Hehuanmou, 2-X-86 (female HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (10 males PARATYPUS, 5 females PARATYPUS in col. T. Mizunuma, Japan).

Taiwan : Nantou Distr. : Mt. Hehuanmou, 24-IX-83 (2 males PARATYPUS in col. H. Fujita, Japan).

Taiwan : Nantou Distr. : Mt. Hehuanmou, 22/29-IX-84 (5 males PARATYPUS, 3 females PARATYPUS in col. M. Baba, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus eugeniae BOMANS, 1991.

Neolucanus eugeniae BOMANS, 1991:363-365.

Neolucanus eugeniae BOMANS; Mizunuma & Nagai, 1994:11, fig. 13(a-b, d); 219, pl. 20.

Sakaino, Yu & Chu, 1994:123.

Taiwan : Kaohsiung Distr. (Liukuei), ? (Rokki Takao*).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus extremus KRIESCHE, 1940.

Kriesche, 1940:40; Benesh, 1960:124; Maes, 1992:28.

China : Yunnan (Schansi--Thai-Yuan-Fu*).

Neolucanus fuscus DIDIER, 1926.

Didier, 1926:213; Didier, 1930:145; Benesh, 1960:124; Maes, 1992:28; Mizunuma & Nagai, 1994:311, pl.149; Baba, 1995:8.

Vietnam : Tonkin (Tuyen-Quang*).

Neolucanus giganteus POUILLAUDE, 1914.

Pouillaude, 1914:37; Benesh, 1960:124; Mizunuma, 1985:15-16, Maes, 1992:28; Mizunuma & Nagai, 1994:223, pl. 23; Kawano, 1997:454.

S. China;

Thailand : Chiang Mai (Chiang Mai--18°50N-98°57E, Nan);

Laos : Samneua;

Vietnam : Tonkin (Dong-Van*, Tam Dao near Vinh Yen).

Neolucanus guiardi DIDIER, 1926.

Didier, 1926:212; Didier, 1929:104; Benesh, 1960:124; Maes, 1992:28; Mizunuma & Nagai, 1994:220, 311, pl.149.

Laos : Phong-Salg*, Xieng-Khouang*.

Neolucanus imitator KRIESCHE, 1935.

Kriesche, 1935:170; Benesh, 1960:124; Maes, 1992:29.

China : Hunan (Sinning*).

Neolucanus insularis MIWA, 1929 [Calcodes].

Miwa, 1929:73; Miwa, 1932:97; Benesh, 1960:124; Nomura, 1960:43; Nomura, 1966:69; Toba, 1987:42-43; Yamaguchi, 1987:30-31; Isida & Fujioka, 1988:4; Araya, 1989:6; Maes, 1992:29; Mizunuma & Nagai, 1994:219, pl. 20; Araya, 1996:20.

Japan : Okinawa Pref. [Ishigakijima I.*--24°24N-124°10E (Mt. Maesedake), Iriomotejima I.--24°20N-123°46E].

 

Neolucanus insulicola ssp. protogenetivus KUROSAWA, 1976.

Neolucanus saundersi ssp. protogenetivus KUROSAWA, 1976.

Neolucanus protogenetivus ssp. protogenetivus KUROSAWA; Mizunuma & Nagai, 1994:12, fig. 19(a-e).

Japan : Kagoshima Pref. [Amami-oshima I. (Mt. Yuwan-dake, Funcha Pass, Onkachi, Setouchi-chô = Nishikata-son*), Tokunoshima I. (Isen)].

 

Neolucanus insulicola ssp. hamaii MIZUNUMA, 1994.

Neolucanus protogenetivus ssp. hamaii MIZUNUMA in MIZUNUMA & NAGAI, 1994:10, 12, fig. 20(a-e), pl. 138, fig. 141(4) [Mt. Oyama].

Japan : Kagoshima Pref. : Amami Guntou Is. : Ukejima I. : Setouchi Cho : Mt. Oyama, 9/11-IX-92, leg. K. Hamai (male HOLOTYPUS, female ALLOTYPUS in Kanagawa Prefectural Museum, Japan) (8 males PARATYPUS, 2 females PARATYPES in col. T. Mizunuma, Japan) (1 male PARATYPUS in col. H. Fujita).

Japan : Kagoshima Pref. : Amami Guntou Is. : Ukejima I. : Setouchi Cho : Mt. Oyama, 1-IX-92, leg. K. Hamai (1 male PARATYPUS in col. T. Mizunuma, Japan).

Japan : Kagoshima Pref. : Amami Guntou Is. : Ukejima I. : Setouchi Cho : Mt. Oyama, 1-X-93, leg. K. Hamai (2 males PARATYPUS, 3 females PARATYPUS in col. T. Mizunuma, Japan) (1 female PARATYPUS in col. H. Fujita, Japan).

 

Neolucanus insulicola ssp. okinawanus SAKAINO, 1984.

Neolucanus saundersi ssp. okinawanus SAKAINO, 1984.

Japan : Okinawa Pref. [Okinawa I.--26°30N-128°E = Okinawahonto I. (Mt. Yonaha-dake*, Oku-river*, Gaji-rindô*, Benoki-river*, Yona-river*, Yona*, Ié-rindô*, Terukubiyama--Timber Road--Kunigami-son)].

 

Neolucanus insulicola ssp. insulicola KUROSAWA, 1976.

Neolucanus saundersi ssp. insulicola KUROSAWA, 1976.

Japan : Okinawa Pref. [Ishigakijima I.*--24°24N-124°10E (Mt. Bannadake), Iriomotejima I.*--24°20N-123°46E (Otomi)].

 

Neolucanus insulicola ssp. donan MIZUNUMA, 1985.

Neolucanus saundersi ssp. donan MIZUNUMA, 1985.

Japan : Okinawa Pref. [Yonagunijima I.--24°28N-122°57E (Mt. Urabudake*, Higawa*)].

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

 

Neolucanus laticollis (THUNBERG, 1806) [Lucanus].

= Dorcus glabratus (DE HAAN) DEJEAN 1837 [nomen nudum] [Java].

Thunberg, 1806:197; Schonherr, 1817:327; Dejean, 1837:193; Hope & Westwood, 1845:18; Albers, 1884:303; Leuthner, 1885:420, pl.84; Heyne & Taschenberg, 1908:52; Oberthur & Houlbert, 1913:325; Benesh, 1960:124; Maes, 1992:29; Mizunuma & Nagai, 1994:220, pl. 21; Kawano, 1997:454.

Java* : Pengalengan, Toegoe, Mont Gedeh, Mt. Salak, Soekaboemi, Wangoen-Tjisampara, Tjigocka, Tjiangsana, Cibodas, Mt. Mas.

Neolucanus latus BOILEAU, 1902 [Calcodes].

= Neolucanus brevis BOILEAU 1899 (female) [Birmanie : Catcin Cauri].

= Neolucanus apricans MOLLENKAMP 1912 [Naga Hills; Brahmaputra].

Boileau, 1899:197; Boileau, 1902:204; Mollenkamp, 1912:302; Arrow, 1943:135; Arrow, 1950:203, pl.21; Benesh, 1960:124; Bomans, 1991:147; Maes, 1992:29; Mizunuma & Nagai, 1994:220, pl. 21.

India : Assam (Brahmaputra), Nagaland (Naga Hills, Kohima);

Myanmar = Burma : Kachin State (Kachin Kauri), Rangoon State (Rangoon--16°48N-96°08E), Karen State (Carin Cheba*, Cheba-Karen Hills), ? (Thandaung);

Thailand : Chiang Mai (Chiang Mai--18°50N-98°57E, Mt. Doi Inthanon--18°26N-98°17E), Tak (Maesot), Loei (Lomkao near Lom Sak, Dansai--Loei near Lom Sak), Petchaburi (Khao Khor);

Laos.

 

Neolucanus lehmanni BABA, 1995.

Neolucanus lehmanni BABA, 1995:7-11 [Dawna Range].

Neolucanus lehmanni BABA; Araya, 1996:16.

Neolucanus lehmanni BABA; Bomans & Miyashita, 1997:3.

Myanmar = Burma : Karen State (Dawna Range*).

Bibliography :

BOMAN H. & MIYASHITA T. (1997) Description de trois nouvelles espèces de Lucanidae de nord Birmanie. 91 ème contribution à l'étude des Coléoptères Lucanides. Besoiro, 4:2-3.

 

Neolucanus lividus DIDIER, 1930.

Didier, 1930:142; Benesh, 1960:124; Maes, 1992:29; Mizunuma & Nagai, 1994:311, pl.149.

China : Fujian (Fuzhou = Foo-chou*).

Neolucanus maculosus DIDIER, 1930 [Localité inconnue].

Didier, 1930:143; Benesh, 1943:37; Benesh, 1960:124; Maes, 1992:29; Mizunuma & Nagai, 1994:311, pl.149.

Sumatra : ? (Bangkinang*ª);

Java.

ª : Locality of the allotype of Benesh 1943.

Note : Mizunuma & Nagai (1994) given Java as type locality for maculosus.

Neolucanus marginatus WATERHOUSE, 1872 [Calcodes].

Waterhouse, 1872:53; Leuthner, 1885:178; Heyne & Taschenberg, 1908:52, pl.7; Boileau, 1913:247; Houlbert, 1914:265; Gravely, 1915:414; Arrow, 1943:135; Arrow, 1950:196, pl.20; Didier & Séguy, 1953:34; Benesh, 1960:124; De Lisle & Chujo, 1964:179; Bomans & Ratti, 1976:176; Sakaino, 1981:50; Bomans, 1989:5; Maes, 1992:29, 30; Bomans, 1994:186-187; Bomans, 1994:490; Mizunuma & Nagai, 1994:220, pl. 21.

Neolucanus marginatus ssp. marginatus WATERHOUSE, 1872.

India* : ? Sikkim (Lacken), Assam, Meghalaya (Khasi Hills), Nagaland (Naga Hills), Manipur;

China : Yunnan (Sse-Tsong).

Neolucanus marginatus ssp. similis BOMANS & RATTI, 1976.

Laos : Ban Van Heua*, P.K.Khouai* (Vientiane), Parkading*, Thong Pheng*, Sayabouri*.

Neolucanus marginatus ssp. dohertyi HOULBERT, 1914.

Myanmar = Burma : Mon State (Kyaikkami = Amherst--Misty Hollow to Sukli), ? (Ruby Mines*, Sima, Kambaiti);

N. Vietnam : Tonkin (Chapa).

 

Neolucanus maximus ssp. confucius LACROIX, 1972.

Neolucanus confucius LACROIX, 1972.

Neolucanus maximus ssp. confucius LACROIX; Mizunuma & Nagai, 1994:11, fig. 16(a, c-e).

India : Darjeeling, Assam, Meghalaya (Shillong*);

Bangladesh;

Thailand : Chiang Rai (Chiang Rai).

 

Neolucanus maximus ssp. fujitai MIZUNUMA, 1994.

Neolucanus maximus ssp. fujitai MIZUNUMA in MIZUNUMA & NAGAI, 1994:10-11, fig. 15(a, c), pl. 22, fig. 139(7), pl. 138, fig. 139(8-10) [Mts. Wuyi Shan, Longyan].

China : Fujian : Wuyi Shan Mts., VIII-1986 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan)

China : Fujian : Wuyi Shan Mts., VIII/IX-1992 (female ALLOTYPUS in Kanagawa Prefectural Museum, Japan) (3 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma) (2 males PARATYPUS, 1 female PARATYPUS in col. H. Yasuoka) (1 male PARATYPUS, 1 female PARATYPUS in col. H. Fujita).

China : Fujian : Wuyi Shan Mts., VIII-1991 (2 males PARATYPUS in col. T. Mizunuma).

China : Fujian : Longyan, VIII/IX-1992 (3 males PARATYPUS, 2 females PARATYPUS in col. T. Mizunuma).

 

Neolucanus maximus ssp. maximus HOULBERT, 1912.

Neolucanus maximus HOULBERT, 1912.

China : Xizang = Thibet (Lou-Tse-Kiang*);

Thailand : Chiang Rai (Wiangpapao), Chiang Mai (Mt. Doi Pui, Fang);

Laos;

N. Vietnam : Tonkin (Sapa--Hoang Lien So'n).

 

Neolucanus maximus ssp. vendli DUDICH, 1923.

Neolucanus vendli DUDICH, 1923.

Taiwan : Taoyuan Distr. (Mt. Rara Shan), Taichung Distr., Nantou Distr. (Nanshanhsi), Kaohsiung Distr. (Liukuei), ? (Candidius Lake*).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus montanus KRIESCHE, 1935.

Kriesche, 1935:169; Benesh, 1960:124; Maes, 1992:29.

China : Sichuan = Ssetschuan (Kinfuschan Mts.--Sung-kan-ho*).

 

Neolucanus nitidus ssp. hengshanensis ICHIKAWA & FUJITA, 1987.

Neolucanus nitidus ssp. hengshanensis ICHIKAWA & FUJITA, 1987.

China : Hunan (Hengshan*, Mt. Huang Shan).

 

Neolucanus nitidus ssp. nitidus (SAUNDERS, 1854) [Odontolabris, Odontolabis].

Neolucanus nitidus ssp. nitidus (SAUNDERS, 1854) [Odontolabris, Odontolabis].

Saunders, 1854:47, pl.4; Leuthner, 1885:427; Boileau, 1913:246; Boileau, 1914:133; Houlbert, 1914:260; Didier, 1926:86; Didier, 1926:210; Didier, 1928:53; Didier, 1929:101; Benesh, 1932:41; Arrow, 1950:199, pl.21; Didier & Séguy, 1953:92; Benesh, 1960:125; Bomans, 1970:226; Ichikawa & Fujita, 1987:12-13; Lacroix, 1988:6; Bomans, 1989:5; Bomans, 1991:147; Maes, 1992:29, 30; Mizunuma & Nagai, 1994:8, 218-219, 311, pl. 19, 149; Kawano, 1997:454.

China* : ? Shanghai (Shanghai), Sichuan, Yunnan, Guangxi, Jiangxi, Zhejiang (Mt. Tianmu Shan), Guangdong (Mt. Loufu Shan), Fujian (Mt. Wuyi Shan, Kuatun).

 

Neolucanus nitidus ssp. hainanensis MIZUNUMA, 1994.

Neolucanus nitidus ssp. hainanensis MIZUNUMA in MIZUNUMA & NAGAI, 1994:8, pl. 19, fig. 126(15-17).

China : Hainan I. : Mt. Wuzhi Shan, 17-VIII-89 (male HOLOTYPUS, female ALLOTYPUS in col. Y. Nose).

China : Hainan I. : Mt. Wuzhi Shan, 26-VIII-89 (1 male PARATYPUS in col. Y. Nose).

 

Neolucanus nitidus ssp. robustus BOILEAU, 1914.

Neolucanus nitidus ssp. robustus BOILEAU, 1914.

= Neolucanus lemeei HOULBERT 1914 [Tonkin] (tal vez es buena especie).

India : Assam;

Myanmar = Burma : South Shan State (Loimwe);

Laos* : Ban Van Heua, Muong-Soui, Thonpheng, P.K. Khouei (Vientiane), Xieng Khouang, Phou Kao;

Vietnam : Tonkin (Hoa-Binh, Ngan-Son, Tam Dao near Vinh Yen).

Note : Bomans (1991) in his work on Lucanidae from Thailand use N. lemeei (from Chiang Mai--18°50N-98°57E) and N. robustus. It seems that    there are two different species.

 

Neolucanus nitidus ssp. maekajanensis ICHIKAWA & FUJITA, 1987.

Neolucanus nitidus ssp. maekajanensis ICHIKAWA & FUJITA, 1987.

Thailand : Chiang Rai (Maekajan*, Wiangpapao); Chiang Mai (Sankampaeng).

Neolucanus nitidus ssp. atratus DIDIER 1926.

Laos;

N. Vietnam : Tonkin (Lao Kay*, Chapa*, Sapa, Sapa--Hoang Lien So'n).

Note : MIZUNUMA & NAGAI (1994) given Laos as type locality for atratus.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus oberthuri ssp. oberthuri LEUTHNER, 1885.

Neolucanus oberthuri LEUTHNER, 1885.

Leuthner, 1885:429; Houlbert, 1914:252, 254; Didier, 1929:15; Benesh, 1960:125; Bomans, 1991:146; Maes, 1992:29, Mizunuma & Nagai, 1994:9, 218, pl. 18.

Neolucanus sinicus ssp. oberthueri LEUTHNER; Mizunuma & Nagai, 1994:9, fig. 10(a-b, d).

Note : Mizunuma & Nagai (1994) consider this species as a ssp. of N. sinicus.

China : SE. Yunnan (Sse-Tsong), Guizhou = Kouy Tcheou*.

 

Neolucanus oberthuri ssp. bisignatus HOULBERT, 1914.

Neolucanus bisignatus HOULBERT, 1914.

Thailand : Chiang Mai (Mt. Doi Inthanon--18°26N-98°17E);

Vietnam : Tonkin (Bao-Lac*).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus oxyops DE LISLE, 1976.

De Lisle, 1976:273-274; Maes, 1992:29.

India : Assam*.

Neolucanus pallescens LEUTHNER, 1885.

Leuthner, 1885:426, pl.85; Benesh, 1960:125; De Lisle, 1976:273; Maes, 1992:30.

China* : Yunnan (Tchaotiong), Fujian, Hong Kong.

Neolucanus palmatus DIDIER & SEGUY, 1952.

Didier & Séguy, 1952:223; Didier & Séguy, 1953:34; Benesh, 1960:125; Maes, 1992:30; Mizunuma & Nagai, 1994:311, pl.149.

Vietnam : Tonkin*.

Neolucanus parryi LEUTHNER, 1885.

= Neolucanus leuthneri BOILEAU 1899 [Tonkin : Tuyen-Kan].

Leuthner, 1885:424, pl.85; Boileau, 1899:175; Houlbert, 1903; Arrow, 1943:134; Arrow, 1950:196, pl.20; Benesh, 1960:125; De Lisle & Chujo, 1964:179; Bomans, 1970:225; Bomans & Ratti, 1976:176; Sakaino, 1981:50-51; Klausnitzer, 1982:43; Kawano, 1988:22; Kawano, 1989:17; Bomans, 1991:147; Maes, 1992:30; Kawano, 1992:12; Mizunuma & Nagai, 1994:220, pl. 21; Kawano, 1997:454, 457, fig. 4.

Myanmar = Burma : Karen State (Cheba-Karen Hills);

China : Guizhou = Kouy Tcheou*;

Thailand = Siam* : Chiang Rai (Wiangpapao, Maekajan), Chiang Mai (Mt. Doi Pui, Mt. Doi Inthanon--18°26N-98°17E, Maetaeng, Sankampaeng, Chiang Mai--18°50N-98°57E, Doi Suthep);

Laos : Pakse, Tonpheng, Sayabouri, P.K. Khouei, Ban Van Heua;

Cambodia;

Vietnam : Tonkin (Rivière Claire affluent Fleuve Rouge, Tuyen-Kan, Bao-Lac, Chiem-Hoa, Tam Dao near Vinh Yen).

Neolucanus perarmatus DIDIER, 1925.

= Neolucanus goral KRIESCHE 1926 [Kouantung : Drachenkopf].

Didier, 1925:262; Kriesche, 1926:362; Didier, 1928:53; Kriesche, 1935:171; Didier & Séguy, 1953:94; Benesh, 1960:125; Bomans, 1970:225; Mizunuma, 1985:15-16; Bomans, 1989:5; Bomans, 1991:147; Maes, 1992:30;        Mizunuma & Nagai, 1994:222, pl. 23; Kawano, 1997:454; Kawano, 1997:25.

China : Guangdong = Kuantung (Drachenkopf), Fujian = Fokien* (Mt. Wuyi Shan, Kuatun);

Thailand : Chiang Rai (Wiangpapao, Maekajan);

Laos* : P.K.Khouei, Ban Van Heua;

Vietnam : Tonkin (Tam Dao near Vinh Yen).

 

Neolucanus rufus NAGEL, 1941.

Nagel, 1941:55; Benesh, 1960:125; Maes, 1992:30.

Vietnam : Tonkin (Chapa*).

Neolucanus rutilans BOMANS, 1989.

Bomans, 1989:5, 14-15; Maes, 1992:30.

China : Fujian (Kuatun*).

Neolucanus sarrauti HOULBERT, 1912.

Houlbert, 1912:183-188; Vitalis, 1919:179; Arrow, 1943:134; Didier & Séguy, 1953:96; Benesh, 1960:123; Lacroix, 1972:122-125; Mizunuma & Nagai, 1994:8-9, 217-218, pl. 18, 138.

Cambodia : Pnom-Penh*;

Vietnam : Tonkin (Tam Dao, Tam Dao near Vinh Yen).

 

Neolucanus saundersi PARRY, 1864.

Calcodes saundersi PARRY, 1864:20, pl. 9.

Leuthner, 1885:431, pl.85; Houlbert, 1912:193; Pouillaude, 1914:40; Dudich, 1923:17; Miwa, 1929:72; Miwa, 1932:94; Arrow, 1950:205, pl.18; Didier & Séguy, 1953:35; Benesh, 1960:122, 125; Nomura, 1966:69; Lacroix, 1972:114, 116; Kurosawa, 1976:190; Lacroix, 1981:64; Sakaino, 1984:8-12; Mizunuma, 1985:15-23; Ito, 1987:36; Isida & Fujioka, 1988:4; Kawano, 1988:22; Araya, 1989:6; Araya, Obuchi & Ochi, 1991:23; Bomans, 1991:147; Araya & Obuchi, 1992:33; Araya, Obuchi & Ochi, 1992:33; Maes, 1992:29-31; Suzuki, 1992:20-23; Sakaino, Yu & Chu, 1994:123.

Neolucanus saundersii PARRY; Mizunuma & Nagai, 1994:10-11, fig. 17(a, c-e), 220-222, pl. 22-23, fig. 138.

Shimoji, 1995:29-31; Araya, 1996:18, 20, 21; Kawano, 1997:454; Kawano, 1997:25.

India* : West Bengal (Darjeeling, Debrepani), Assam;

Bhutan.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

 

Neolucanus sianoukei LACROIX, 1978.

Lacroix, 1978:269; Maes, 1992:30.

Note : Pierre Benoit (in litteris) consider this as a synonym of Neolucanus delicatus DIDIER. It’s a bicolor medium sized form.

Vietnam : Tonkin (Hanoï*).

 

Neolucanus sinicus ssp. ?

Sikkim.

 

Neolucanus sinicus ssp. sinicus (SAUNDERS, 1854) [Odontolabris].

Odontolabris sinicus SAUNDERS, 1854: 48, pl.4.

Parry, 1864:20; Leuthner, 1885:428, pl.85; Boileau, 1899:176; Heyne & Taschenberg, 1908:52; Houlbert, 1912:83, 183; Houlbert, 1914:254, 257; Didier, 1928:53; Miwa, 1932:97; Arrow, 1943:134; Didier & Séguy, 1953:34; Benesh, 1960:123, 125; Lacroix, 1972:122; Sakaino, 1981:50; Klausnitzer, 1982:45; Bomans, 1989:5; Bomans, 1991:147, 149; Bomans, 1991:363; Maes, 1992:28, 29, 30; Ma, 1993:279; Sakaino, Yu & Chu, 1994:123; Araya, 1996:15, 21; Kawano, 1997:454.

Neolucanus sinicus ssp. sinicus (SAUNDERS); MIZUNUMA & NAGAI, 1994:9, fig. 5(a-d), pp. 217-218, pl. 18, 138.

China* : Sichuan (Miyi), Hunan (Wuling Mts.), Yunnan (Pe-Yen-Tsing, Kunming), Shanghai (Shanghai), Jiangxi (Mt. Zhenzhu Shan), Zhejiang (Pomasishi, Mt. Siming Shan, Mt. Simiang Shan), Fujian = Fokien (Kuatun).

 

Neolucanus sinicus ssp. championi PARRY, 1864.

Neolucanus sinicus ssp. championi PARRY, 1864.

Neolucanus sinicus ssp. championi PARRY; Mizunuma & Nagai, 1994:9, fig. 7(a, c-d).

China* : Guangdong (Mt. Loufu Shan), Hong Kong* (Victoria Peak, Mt. Taimou Shan).

 

Neolucanus sinicus ssp. nosei MIZUNUMA, 1994.

Neolucanus sinicus ssp. nosei MIZUNUMA in MIZUNUMA & NAGAI, 1994:8-9, fig. 8(a-d), pl. 18, fig. 124(19-23).

China : Hainan I. : Mt. Wuzhi Shan, 30-V-1989 (male HOLOTYPUS in col. Y. Nose, Japan).

China : Hainan I. : Mt. Wuzhi Shan, 29-V-1989 (female ALLOTYPUS in col. Y. Nose, Japan).

China : Hainan I. : Mt. Wuzhi Shan, 21-V-1989 (4 males PARATYPUS in col. T. Mizunuma, Japan).

China : Hainan I. : Mt. Wuzhi Shan, 3-VI-1989 (1 male PARATYPUS in col. Y. Nose, Japan).

 

Neolucanus sinicus ssp. taiwanus MIZUNUMA, 1994.

Neolucanus sinicus ssp. taiwanus MIZUNUMA in MIZUNUMA & NAGAI, 1994:8-9, fig. 5(b-c); lam. 18 fig. 124(17-18) [Hungye].

Taiwan : Hualien Distr. : Juisui : Hungye, 800-1000 m, 10/20-VI-1973 (male HOLOTYPUS in Kanagawa Prefectural Museum, Odawara, Japan) (1 male Paratypus in col. T. Mizunuma).

Taiwan : Nantou Distr. : Puli.

Taiwan : ? Hengschun.

 

Neolucanus sinicus ssp. fuliginatus MIZUNUMA, 1994.

Neolucanus sinicus ssp. fuliginatus MIZUNUMA in MIZUNUMA & NAGAI, 1994:8-9, fig. 11a [Jinghong].

Neolucanus sinicus ssp. fulginatus MIZUNUMA; MIZUNUMA & NAGAI, 1994:pl. 18, figs. 124(32).

China : S. Yunnan : Jinghong, VII-1991 (male HOLOTYPUS in col. Kanagawa Prefectural Museum, Odawara, Japan) (4 males PARATYPUS in col. T. Mizunuma) (1 male PARATYPUS in col. H. Fujita).

 

Neolucanus sinicus ssp. pseudopacus HOULBERT, 1914.

Neolucanus sinicus ssp. pseudopacus HOULBERT, 1914.

= Neolucanus intermedius HOULBERT 1914 [Kouy-Tcheou].

Myanmar = Burma;

China : Guizhou = Kouy Tcheou*;

Thailand : Chiang Mai (Mt. Doi Inthanon--18°26N-98°17E), Petchaburi (Khao Khor).

 

Neolocanus sinicus ssp. opacus BOILEAU, 1899.

Neolocanus sinicus ssp. opacus BOILEAU, 1899.

Neolocanus sinicus ssp. opacus BOILEAU; Mizunuma & Nagai, 1994:9, fig. 9(a, c-d).

Cambodia;

N. Vietnam : Tonkin (Kiu-Kiang*--Po-Yang Lake, Hoa-Binh, Than-Moï, Chapa--Lao-Kay, Mt. Manson, Bas Tonkin-Delta, Tam Dao near Vinh Yen, Sapa--Hong Lien So'n);

Malaya.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus spicatus DIDIER, 1930.

Didier, 1930:141; Benesh, 1960:125; Maes, 1992:30; Mizunuma & Nagai, 1994:311, pl.149.

China : Fujian (Foochou* = Fou-Tcheou = Foochow).

 

Neolucanus swinhoei BATES, 1866.

Neolucanus swinhoei BATES, 1866:346.

Parry, 1870:57; Leuthner, 1885:424, pl.84; Miwa, 1932:95; Didier & Séguy, 1953:59; Benesh, 1960:125; Klausnitzer, 1982:43;           Ichikawa & Fujita, 1987:12-13; Maes, 1992:30.

Neolucanus swinhoei BATES; Mizunuma & Nagai, 1994:11, fig. 14(a, c-d); 219, pl. 20.

Sakaino, Yu & Chu, 1994:123; Kawano, 1997:25.

Taiwan* : Taoyuan Distr. (Mt. Rara Shan), Taipei Distr. (Mt. Tatun Shan), Nantou Distr. (Wushai, Puli), Kaohsiung Distr. (Liukuei), Taitung Distr. (Chihpen wenchuan), ? (Pingtung, Chip Chip, Hoozan, Kosempo, Suisharyo, Hokuto).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus tanakai MIZUNUMA, 1994.

Neolucanus tanakai MIZUNUMA in MIZUNUMA & NAGAI, 1994:10-11, fig. 18(a-e), pl. 23, fig. 144(1) [Mt. Wuzhi Shan].

China : Hainan I. : Mt. Wuzhi Shan, 1800 m, 20-VIII-89 (male HOLOTYPUS in Kanagawa Prefectural Museum).

China : Hainan I. : Mt. Wuzhi Shan, 1800 m, 19-VIII-89 (1 male PARATYPUS in col. Masahiro Tanaka, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Neolucanus tao KRIESCHE, 1935 [taos].

Kriesche, 1935:170; Benesh, 1960:125; Maes, 1992:30.

China : Guangxi = Kwangsi (Toyen-chan Mts.*).

Neolucanus vicinus POUILLAUDE, 1913.

Pouillaude, 1913:333; Arrow, 1943:135; Didier & Séguy, 1953:96; Benesh, 1960:125; Maes, 1992:31; Mizunuma & Nagai, 1994:219, pl. 20.

Vietnam : Tonkin* (Chapa--Lao-kay, Sapa--Hong Lien So'n).

Neolucanus zebra LACROIX, 1988.

Lacroix, 1988:5-6; Araya, 1996:22.

Taiwan : Nantou Distr. (Puli = Puly*).


HOME

LUCANIDAE

GENERA