Cyclommatus imperator

Cyclommatus (Cyclommatus) imperator BOILEAU, 1905.

Cyclommatus (Cyclommatus) imperator ssp. monguilloni LACROIX, 1981.

Cyclommatus monguilloni LACROIX, 1981:55-56, fig. [Fak Fak].

Cyclommatus imperator monguilloni LACROIX; Mizunuma & Nagai, 1991:8.

Cyclommatus monguilloni LACROIX; Maes, 1992:59.

Cyclommatus imperator monguilloni LACROIX; Mizunuma & Nagai, 1994:236, pl. 48, fig.224(9-15), pl.138, fig.224(8).

Cyclommatus monguilloni LACROIX; Taroni, 1998:32, 57, fig.

Cyclommatus monguilloni LACROIX; Schenk, 2000:10-11.

Cyclommatus imperator monguilloni LACROIX; Krajcik, 2001:32.

Ø New Guinea : Irian Jaya : Fak Fak, 2°55S-132°15E, 19-XI-1978, Monguillon leg. (male HOLOTYPUS of Cyclommatus monguillonni LACROIX, 1981).

Ø New Guinea : Irian Jaya : Fak Fak, 2°55S-132°15E, XI-1980, G. Lachaume leg. (male PARATYPUS of Cyclommatus monguillonni LACROIX, 1981).

Ø New Guinea : Irian Jaya : Mt. Arfak

Ø New Guinea : Irian Jaya : Mt. Arfak : Cendrawasih, V-1990 (1 male in coll. Mizunuma, Japan).

Ø New Guinea : Irian Jaya : Mt. Ilaga

Ø New Guinea : Irian Jaya : Pegunungan Maoke : Mt. Ilaga, III-1990 (2 males in coll. Mizunuma, Japan).

Ø New Guinea : Irian Jaya : Pegunungan Maoke : Mt. Ilaga, II-1991 (2 males in coll. Mizunuma, Japan).

Ø New Guinea : Irian Jaya : Mapia, IV-1992 (1 male in coll. Mizunuma, Japan).

Ø New Guinea : Irian Jaya : Mapia, VI-1992 (1 male in coll. Mizunuma, Japan).

Ø New Guinea : Irian Jaya (1 male in coll. Taroni, Italy).


TYPUS (adapted from Lacroix).	Adapted from Mizunuma & Nagai.

Adapted from Mizunuma & Nagai.		Adapted from Taroni.

Bibliography :

KRAJCIK M. (2001) Lucanidae of the World. Catalogue – Part.I. Checklist of the Stag Beetles of the World (Coleoptera: Lucanidae). Czech Republic, 108 pp.

LACROIX J.P. (1981) Description d'un Cyclommatus nouveau (Coleoptera Lucanidae) de la faune néo-guinéenne. Bulletin de la Société Sciences Nat 29-30: 55-56.

MAES J.M. (1992) Lista de los Lucanidae (Coleoptera) del Mundo. Revista Nicaraguense de Entomología, 22:121 pp.

MIZUNUMA T. & NAGAI S. (1991) A new species and four new subspecies of the genus Cyclommatus (Coleoptera, Lucanidae). Gekkan-Mushi, 244:4-10.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

SCHENK K.D. (2000) Beschreibung einer neue Art der Gattung Cyclommatus Parry, 1863, aus West-Neuguinea (Coleoptera, Lucanidae). Entomologische Zeitschrift, 110(1):10-11.

TARONI G. (1998) Il Cervo Volante (Coleoptera Lucanidae). Electra, 182 pp.

Cyclommatus (Cyclommatus) imperator ssp. imperator BOILEAU, 1905.

Cyclommatus imperator BOILEAU, 1905:285, fig. (Owgarra).

Cyclommatus imperator BOILEAU; Van Roon, 1910:29.

Cyclommatus imperator BOILEAU; Didier, 1930:137, 139-140.

Cyclommatus imperator BOILEAU; Nagel, 1932:82-83, 85.

Cyclommatus imperator BOILEAU; Nagel, 1932: 114.

Cyclommatus imperator BOILEAU; Didier & Séguy, 1952:pl.86, figs.1-8.

Cyclommatus imperator BOILEAU; Didier & Séguy, 1953:68, 122.

Cycommatus imperator var. ochracea NAGEL; Didier & Seguy, 1953:15, 122, fig.1 [Sattelberg] (unpublished name; Didier & Séguy said Chirotype).

Cyclommatus imperator BOILEAU; Benesh, 1960:58.

Cyclommatus imperator BOILEAU; De Lisle, 1968:399.

Cyclommatus imperator BOILEAU; Lacroix, 1972:58-59.

Cyclommatus imperator BOILEAU; De Lisle, 1980:286.

Cyclommatus imperator BOILEAU; Lacroix, 1981:55-56.

Lacroix, Ratti & Taroni, 1983:6

Bomans, 1984:354

Bomans, 1988:6

Cyclommatus imperator BOILEAU; Mizunuma & Nagai, 1991:8.

Ratcliffe, 1991:252

Cyclommatus (Cyclommatus) imperator BOILEAU; Maes, 1992:58.

Cyclommatus imperator imperator BOILEAU; Mizunuma & Nagai, 1994:236, pl. 48, fig.224(1-8).

Kawano, 1997:25.

Cyclommatus imperator BOILEAU; Taroni, 1998:32, 57, figs.

Cyclommatus imperator BOILEAU; Emden & Nijhout, 2000:662, fig.

Cyclommatus imperator BOILEAU; Schenk, 2000:10-11.

Cyclommatus imperator BOILEAU; Krajcik, 2001:32.

Cyclommatus imperator BOILEAU; Krajcik, 2003:51.

Ø New Guinea : Irian Jaya : Wamena, III-1993 (1 male in coll. Mizunuma, Japan).

Ø New Guinea : Papua : Pausa, 1700 m, II-1979, J. Sedlacek leg., ex coll. Institut royal des Sciences naturelles de Belgique (1 male in coll. Museo Entomológico de León, Nicaragua).

Ø New Guinea : Papua : Pausa, 1700 m, II-1979, J. Sedlacek leg. (1 male, 1 female in coll. Institut royal des Sciences naturelles de Belgique, Belgium).

Ø New Guinea : Papua : Bismarck Range : Okapa EHP, V-1982, leg. G. Debatisse (2 males in coll. Museo Entomologico de Leon, Nicaragua).

Ø New Guinea : Papua : Bismarck Range : Okapa, XII-1985 (1 male in coll. Mizunuma, Japan).

Ø New Guinea : Papua : Bismarck Range : Okapa, V-1989 (2 males in coll. Mizunuma, Japan).

Ø New Guinea : Papua : Bismarck Range : Okapa, VIII-1989 (1 male in coll. Mizunuma, Japan).

Ø New Guinea : Papua : Okapa EHP, V-1982, ex coll. G. Debatisse (2 males in coll. Museo Entomológico de León).

Ø New Guinea : Papua : Owen Stanley Range : Bulolo : Yamap, V-1989 (2 males in coll. Mizunuma, Japan).

Ø New Guinea : Papua : Bismarck Range : Jimmi, V-1989 (1 female in coll. Mizunuma, Japan).

Ø New Guinea : Papua : Sattelberg (male TYPUS of Cycommatus imperator var. ochracea NAGEL; Didier & Seguy, 1953, undescribed name).

Ø New Guinea : Papua : Goroka

Ø New Guinea : Papua : Eastern Highlands : Okapa

Ø New Guinea : Papua : Owgarra = Ugara = Ougara, A.S. Meek leg. (male HOLOTYPUS and female ALLOTYPUS of Cyclommatus imperator BOILEAU, 1905 in MNHN, Paris).

Ø New Guinea (6 males in coll. Didier, in coll. MNHN, Paris; figured in Didier & Séguy, 1952).

Ø New Guinea (17 males, 4 females in coll. Taroni, Italy).


Adaptado de Didier & Séguy.	Adaptado de Mizunuma & Nagai.		Papua New Guinea : Okapa, leg. G. Debatisse (in coll. Museo Entomologico de Leon, Nicaragua).

Papua New Guinea : Okapa, leg. G. Debatisse (in coll. Museo Entomologico de Leon, Nicaragua).

Papua New Guinea : Pausa, leg. J. Sedlacek (in coll. Museo Entomologico de Leon, Nicaragua).	Adapted from Didider & Séguy.

Alive specimen (Picture J. Oldrich in Ben Harink’s web site)		Alive specimen (Picture J. Oldrich in Ben Harink’s web site)

Alive pair originated from Papua New Guinea : Wamena, hatched August 2005 (Picture Gomi Takahiro)

*Cyclommatus imperator* *ssp. arfakensis* WAKATAKE (adapted from a Japanese web site, can’t understand correct reference).**	*Cyclommatus imperator* *ssp. splendidus* SCHENK (adapted from a Japanese web site, can’t understand correct reference).**

Bibliography:

BENESH B. (1960) Lucanidea. Coleopterorum Catalogus, Supplementa (Second Edition) 8: 1-178.

BOILEAU M.H. (1905) Description de Coléoptères nouveaux. Le Naturaliste 27(451): 285-287.

DE LISLE M.O. (1968) Lucanidae (Col.) récoltés par l'expédition du Noona Dan aux Iles Bismarck et Salomon. Entomologiske Meddelelser 36: 385-405.

DE LISLE M.O. (1980) Note sur les Cyclommatus (Col, Lucanidae) New-guineens du group Margaritae Gestro. Nouvelle Revue d’Entomologie (Nouvelle Série) 10(3): 285-299.

DIDIER R. (1930) Études sur les Coléoptères Lucanides du globe. VIII. Descriptions de Lucanides nouveaux ou peu connus. IX. Études d'une collection de Lucanides de l'Inde. X. Descriptions de Lucanides nouveaux. Librairie speciale Agricole, Paris, fascicule 6: 125-140.

DIDIER R. & SEGUY E. (1952) Catalogue illustré des Lucanides du Globe. Atlas. Encyclopedie Entomologique, Serie A 28: 112 plates.

DIDIER R. & SEGUY E. (1953) Catalogue illustré des Lucanides du Globe. Texte. Encyclopedie Entomologique, Serie A 27: 1-223.

EMLEN D.J. & NIJHOUT H.F. (2000) The development and evolution of exaggerated morphologies in insects. Annual Review of Entomology, 45:661-708.

KRAJCIK M. (2001) Lucanidae of the World. Catalogue – Part.I. Checklist of the Stag Beetles of the World (Coleoptera: Lucanidae). Czech Republic, 108 pp.

KRAJCIK M. (2003) Lucanidae of the World. Catalogue – Part.II. Encyclopaedia of the Lucanidae (Coleoptera: Lucanidae). Czech Republic, 197 pp., 10 plates.

LACROIX J.P. (1972) Descriptions de Coléoptères Lucanidae nouveaux ou peu connus. Bulletin et Annales de la Société royale d’Entomologie de Belgique, 108(1-4): 33-71.