Lucanus (Lucanus) SCOPOLI, 1763

(G. = Scarabaeus cervus)

= Hexaphyllus MULSANT 1839 (G. = Hexaphyllus pontbrianti)


Lucanus (Lucanus) angusticornis DIDIER, 1925.

Lucanus angusticornis (Boileau in litt.) DIDIER, 1925:219 [Laos : Xieng-Khouang].

Lucanus angusticornis DIDIER; Didier, 1926:144

Lucanus angusticornis DIDIER; Didier & Séguy, 1953:81

Lucanus angusticornis DIDIER; Benesh, 1960:131

Lucanus angusticornis DIDIER; Lacroix, 1972:50

Lucanus angusticornis DIDIER; Kurosawa, 1978:143

Lucanus angusticornis DIDIER; Sakaino, 1981:23

Lucanus angusticornis DIDIER; Maes, 1992:15

Lucanus angusticornis DIDIER; Kawano, 1994:9

Lucanus angusticornis DIDIER; Mizunuma & Nagai, 1994:216, pl.15

Lucanus angusticornis DIDIER; Kawano, 1997:454

Lucanus angusticornis DIDIER; Kawano, 1997:25.

Lucanus angusticornis DIDIER; Araya, 2001:10, figs.1-5.

·        Laos : Xieng-Khouang, 18-VI-1919, ex. coll. Boileau (male HOLOTYPE of Lucanus angusticornis [Boileau in litteris] DIDIER, 1925 en Museum National d’Histoire Naturelle, Paris).

·        Vietnam : Tonkin : Tam Dao near Vinh Yen at NE Hanoi, leg. H. Perrot (female TYPE of Lucanus angusticornis DIDIER, 1925 by LACROIX, 1969, published in 1972 en Museum National d’Histoire Naturelle, Paris).

Male HOLOTYPE of Lucanus angusticornis, from Laos (adapted from Araya, 2001).

Female TYPUS of Lucanus angusticornis, from Viet Nam (adapted from Araya, 2001).

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) bruanti LACROIX & BOMANS, 1973.

Lucanus bruanti LACROIX & BOMANS, 1973:138

Lucanus bruanti LACROIX & BOMANS; Mizunuma & Nagai, 1994:309, pl.147.

Lucanus bruanti LACROIX & BOMANS; Araya, 2001:10-11, figs.8-10.

·        India : Assam

·        Bhutan (British Bootan) : Padong. 1914, col. L. Durel, col. R. Oberthur (male HOLOTYPUS of Lucanus bruanti LACROIX & BOMANS, 1973, #667A en Museum National d’Histoire Naturelle, Paris)

Male HOLOTYPUS of Lucanus bruanti LACROIX & BOMANS, 1973, from Bhutan (adapted from Araya, 2001).

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) cambodiensis DIDIER, 1925.

Lucanus cambodiensis DIDIER, 1925:222 [Cambodia : Lao Kay].

Lucanus cambodiensis DIDIER; Didier, 1926:169

Lucanus cambodiensis DIDIER; Didier & Séguy, 1952:pl.17

Lucanus cambodiensis DIDIER; Benesh, 1960:131

Lucanus cambodiensis DIDIER; Maes, 1992:15.

Lucanus cambodiensis DIDIER; Araya, 2001:10-11, figs.11-12.

·        Cambodia : Lao Kay, IX-1919, col. H. Deyrolle (female TYPE of Lucanus cambodiensis DIDIER, 1925 in Museum National d’Histoire Naturelle, Paris).

Female TYPE of Lucanus cambodiensis DIDIER, 1925, from Cambodia (adapted from Araya, 2001).

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

 

Lucanus (Lucanus) cantori ssp. cantori HOPE, 1842 [Khasi Hills].

Lucanus cantori HOPE, 1842:83 [Khasi Hills].

Lucanus cantori HOPE; Hope, 1843:363

Lucanus cantori HOPE; Hope, 1845:73

Lucanus cantori HOPE; Hope & Westwood, 1845:5

Lucanus cantori HOPE; Burmeister, 1847:527

Lucanus cantori HOPE; Planet, 1899:275

Lucanus cantori HOPE; Planet, 1902:57, pl.13

Lucanus cantori HOPE; Heyne & Taschenberg, 1908:50, pl.6

Lucanus cantori HOPE; Gravely, 1915:411

Lucanus cantori HOPE; Didier, 1949:pl.12

Lucanus cantori HOPE; Arrow, 1950:51, pl.3

Lucanus cantori HOPE; Benesh, 1960:131

Lucanus cantori HOPE; Lacroix, 1967:97-98

Lucanus cantori HOPE; Maes, 1992:15-16

Lucanus cantori HOPE; Mizunuma & Nagai, 1994:213, pl.10.

Lucanus cantori HOPE; Araya, 2001:11, fig.13.

·        India : West Bengal : Darjeeling

·        India : West Bengal : Kurseong

·        India : West Bengal : Gopaldhara

·        India : West Bengal : Rungbong Valley

·        India : West Bengal : Darjeeling : Redong

·        India : “?Himalaya”or “Khasi Hills”  (male TYPE of Lucanus cantori HOPE, 1842, # 356 in Oxford University Museum of Natural History).

·        Bhutan : Pedong

·        Bhutan : Paro Distr.

·        Bhutan : Gedu.

Adults on ? : Castanopsis, Symplocos.

Male TYPE of Lucanus cantori HOPE, 1842 (adapted from Araya, 2001).

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) cantori ssp. colasi LACROIX, 1967.

Lucanus cantori ssp. colasi LACROIX, 1967:97-98 [India : Shillong].

·        India : Meghalaya : Shillong (male TYPE of Lucanus cantori ssp. colasi LACROIX, 1967).

·        India : Meghalaya : Khasi Hills.

Note : it seems that the nominal subspecies was describe from Khasi Hills, if it is so, colasi will be synonym of cantori and then the ssp. of Darjeeling is unnamed.

 

Lucanus (Lucanus) cervus ssp. cervus (LINNAEUS, 1758).

Scarabaeus cornutus DALE, 1693:389, pl.17.

Linnaeus, 1745:69; Rossel, 1746:25, pl.4-5

Scarabaeus cervus LINNAEUS, 1758:353.

Poda, 1761:21; Sulzer, 1761:3, pl.1; Geoffroy, 1762:61, pl.1; Pontopp., 1763:430; Kurz., 1765:199; Scopoli, 1763:1; De Bomare, 1764:486; Gronov, 1764:147; Muller, 1764:2; Schaeffer, 1766:pl.9; Linnaeus, 1767:559; Schluga, 1767:pl.1; De Geer, 1774:327, pl.12; Fabricius, 1775:1

Lucanus capreolus FUESSLY, 1775:3

Lucanus dorcas MULLER, 1776:52

Sulzer, 1776:19, pl.2; Bergstrasser, 1778:18, 25, pl.4, 13; Fuessly, 1778:63; Schaeffer, (1766)-1779:pl.133; Barbut, 1781:15, pl.2; Fabricius, 1781:1; Fuessly, 1781:2; Schrank, 1781:32; Von Laichart., 1781:1; Von Scheven, 1782:60; Panzer, 1783:36, pl.29

Lucanus cervus LINNAEUS; De Asso, 1784

Leske, 1784:464; Bonsd., 1785:221; Fourcroy, 1785:2; Von Moll, 1785:149; Fabricius, 1787:1; Petagna, 1787:1; Von Meidinger, 1787:117; Zschach, 1788:1; De Villers, 1789:41

Lucanus capra OLIVIER, 1789:9-12, pl.1-2

Schrank, 1789:374; Von Razoum., 1789:138; Brahm, 1790:124, 130

Lucanus armiger HERBST, 1790:287, 289, 299, 301, pl.D-E, pl.33-34.

Lucanus hircus HERBST, 1790:287, 289, 299, 301, pl.D-E, pl.33-34.

Linnaeus (ed. Gmelin), 1790:1588; Preyssler, 1790:9; Roesel, 1790:1749, pl.iv; Rossi, 1790:1-2; Scriba, 1790:40; Mayer, 1791:181; Donovan, 1792:31, pl.13; Fabricius, 1792:236-237; Kugelann, 1792:255; Martyn, 1792:pl.5; Brahm, 1793:657; Fabricius, 1793:236; Schwarz, 1793:84; Panzer, 1795:244; Hoppe, 1796:169, 179; Hoppe, 1797:179, 200; Panzer, 1798:11;

Olivier (ed. Illiger), 1800:55-59; Paykull, 1800:45; Walther, 1800:352; Fabricius, 1801:248-249

Lucanus inermis MARSHAM, 1802:46, 48, pl.2 [Loc. ?].

Blumenb., 1803:321; Latreille, 1804:246, pl.86; Posselt, 1804:15, pl.ii; Duftschm., 1805:64; Panzer, 1805:110; Bechst. & Scharfenb., 1805:80; Sturm, 1805:pl.1-2; Thunberg, 1806:190-191; Fallen, 1807:12, 22

Lucanus grandis HAWORTH, 1807:77

Latreille, 1807:135; Gyllenhall, 1808:65-66; Schonherr, 1817:318, 320-321; W.S. Macleay, 1819:114; Blot, 1824:84; Muller, 1825:553-672; Klug, 1829:365, pl.15; Stephens, 1830:166; Davis, 1832:86; Curtis, 1834:490; Audoin, 1835:81; Pictet, 1836:393; Dejean, 1837:193; Mulsant, 1839:119, pl.2; Newport, 1839:853-994; Ratzeburg, 1839:105; Westwood, 1839:187; Castelnau, 1840:171; Spry & Shuckard, 1840:34, pl.42; Heer, (1838)-1842:495

Lucanus microcephalus MULSANT, 1842:8, 583, 586, pl.3

La Ferté Senectère en Mulsant, 1842:8; Burmeister, (1835)-1843:pl.24; Sturm, 1843:347, pl.5; Tulk, 1843:451; Brulle, 1844:271-374; Desmarest, 1845:53

Lucanus americanus HOPE & WESTWOOD, 1845:9-10 [N. America, error]

Lucanus lusitanicus HOPE & WESTWOOD, 1845:9-10 [Portugal].

Lucanus vicinus HOPE & WESTWOOD, 1845:9-10 [Poonah, error].

Motschulsky, 1845:60; Bellier de la Chavignerie, 1846:28; Blanchard, 1846:322

Lucanus major BURMEISTER, 1847:350, 527

Erichson, 1848:936, 938; Kriechbaun., 1848:208;

Reiche, 1853:71; Perris, 1854:105; Mulsant & Godart, 1855:250; Mulsant & Godart, 1855:150; Lacordaire, 1856:23; Reiche, 1856:81; Delarouzee, 1859:98; Delarouzee, 1859:42; Duval, 1859:7, 9-10, pl.1; Gerst., 1859:110; Deyrolle, 1860:22; Kraatz, 1860:68, 265, 273, pl.7; Parry, 1864:71; Abeille de Perrin, 1867:65, 70; Cornelius, 1867:435; Cornelius, 1868:33; Landois, 1868:88-95, pl.vi; MacLachlan, 1868:ix; Motschulsky, 1870:25-31, 34, pl.2; De Narcillac, 1871:186; Hochuth, 1871:85; Mulsant & Rey, 1871:8, 14, 16; Schioedte, 1873:342-345, pl.15; Redtenbacher, 1874:451; Schiodte, 1874:341, pl.xviii-xix; Schlechtendal, 1874:25; Schlechtendal, 1875:158; Kraatz, 1876:72; Rossi, 1876:95; Schmidt-Goebel, 1876:392; Perris, 1877:101-102, pl.5; Rossi, 1878:227; Schreiber, 1878:259; Brandt, 1879:1-17; Cotton, 1880:83-84; Reichenau, 1880:55; Rupertsberger, 1880:142; Hammond, 1881:14-26, pl.2; Reichenau, 1881:172-194; Kraatz, 1883:27; Leboeuf, 1883:45; Strubing, 1883:159; Cornelius, 1884:33; Smith, 1885:27; Frivaldsky, 1886:78; Gadeau de Kerville, 1886:176; Fallou, 1889:141; Rousseau, 1889:170; Fowler, 1890:4; Planet, 1890:156-157, 164-165, 215-216

Lucanus antennatus REY, 1890:164

Reitter, 1891:142-143; Verhoeff, 1893:113-170; Verhoeff, 1893:209-260; Binet, 1894:449-580; Pottier, 1894:27; Rupertsberger, 1894:146; Borgschulze, 1895:131; Caradja, 1895:77; Eggers, 1895:131; Schaufuss, 1895:111; Selmons, 1895:52; Planet, 1896:11, 99, 168, 238, 256; Planet, 1897:64, 192, 229, 267; Weise, 1897:389; Bothe, 1899:140; Planet, 1899:31, 35, 42, 46, 55, 58, 62, 64-65, 104, 107, pl.9, 10, 14; Rupertsberger, 1899:235; Stichel, 1899:19

Lucanus longipennis VON ROTHENBURG, 1900:99

Jarder, 1901:28; Le Senechal, 1901:190; Xambeau, 1901:604; Planet, 1902:52, 100, 111, 114-115, 119, 120, 125; Weber, 1902:335; Biedermann, 1903:395-481

Lucanus cervus LINNAEUS; Champion, 1903

Lucanus cervus LINNAEUS; Champion, 1904

Marchand, 1904:37; Rudow, 1904:179, 187, 196; Schuster, 1904:338; Amman, 1905:139; Scabra, 1905:7-8, pl.; Rey, 1906:59; Barthe, 1909:5-6; Chagrin, 1909:404; Reitter, 1909:299, pl.68; Chinaglia, 1910:1-8, 25-39; Leoni, 1910:153

Lucanus capitalis MOLLENKAMP, 1910:242, 575

Schieser, 1910:29; Bedel, 1911:8; Kapzov, 1911:297-337; Krizenecky, 1911:113; Pic, 1911:399; Sasse, 1911:69-104

Lucanus validus MOLLENKAMP, 1912:302 [Borneo, error].

Muzik, 1912:189; Ramme, 1912:37; Roubal, 1912:386; Sirguey, 1912:134; Boileau, 1913:218, 220; Bordas, 1913:358-364; Fairmaire & Planet, 1913:195, pl.9; Kuhnt, 1913:380; Bordas, 1914:181-183; Braatz, (1913)-1914:267; Moritz, 1914:138; Stellwag, 1914:359-429; Chinaglia, 1915:7; Houlbert & Monnot, 1915:4, 6; Labitte, 1916:105; Schaufuss, 1916:1252, pl.48; Verhoeff, 1916:354-372; Della Beffa, 1921:3; Houlbert (1912)-1921:167; Dudich, 1923:129; Grutzner, 1924:130; Nagel, 1924:398-399; Henriksen, 1925:134-143; Nagel, 1928:273

Lucanus ornatus BERNAU, 1929:72

Catherine, 1929:5-6; Vignon, 1929:89-125; Gadeau de Kerville, 1930:63; Nagel, 1930:90; Goldschmidt, 1931

Lucanus nigrinus MARCH, 1931.

Marcu, 1931:214; Portevin, 1931:2-3, pl.6; Roubal, 1931:36; Baudrimont, 1932:116; Franz, 1932:134; Joy, 1932:242; Joy, 1932:pl.69; Kraemer, 1932:321; Kraemer, 1932:86-117; Porta, 1932:371; Didier, 1933:24, pl.1; Hepp, 1933:126-127; Hepp, 1934:84; Golovyanko, 1935:38; Van Emden, 1935:194, 200; Hepp, 1936:173; Paulian, 1936:751-759; Didier, 1937:xxxviii, xlii; Muller & Schoop, 1937:41-116; Palm, 1941:17; Van Emden, 1941:119; Garnier, 1945:110; Sellier & Razet, 1947:223-225; Colas, 1949:128-131; Didier, 1949:pl.1, 3;

Leiler, 1950:157; Muller & Schoop, 1950:17-18; Arrow, 1951:pl.8; Condrillier, 1951:20; Brull, 1952; Didier & Séguy, 1952:pl.15, 17, 24; Tressens, 1952:57; Auber, 1953:99-100; Didier & Séguy, 1953:27-32; Tippmann, 1954; Theodorides, 1955; Britton, 1956:6; Landin, 1956:1; Paulian, 1956:36; Freude, 1957:129-134; Harz, 1957:22-23; Dorn, 1958:62-64

Lucanus cervus LINNAEUS; Español, 1958

Lucanus cervus (LINNAEUS); Horion, 1958

Papperitz, 1958; Weise, 1958; De Ligondes, 1959:52-56; Franz, 1959:74-79; Marie, 1959:125-127; Paulian, 1959:38-39; Weinreich, 1959:282-283; Benesh, 1960:132, 138-140; Janssens, 1960:95; Muspratt, 1960:53; Kuhn, 1961; Colas, 1962:118; Demoulin, 1963:pl.8

Lucanus cervus f. scapulodonta WEINREICH, 1963:29.

Burton, 1965:121; Clark, 1966:199

Lucanus cervus LINNAEUS; Bahuena, 1967

Clark, 1967:24; Lacroix, 1967:98

Lucanus cervus (LINNAEUS); Freude, 1968:59-64

Lacroix, 1968:234-235; Lacroix, 1969:119-129; Machatschke, 1969; Mathieu, 1969:1061; Freude, 1971:283

Lucanus cervus (LINNAEUS); Cofais, 1972:32-33

Lovari, 1972:36

Lucanus cervus (LINNAEUS); Mal, 1972:161-167

Lacroix & Bomans, 1973:152; Kersmaekers, 1973:219; Verstraeten, Boosten & Gaspar, 1973:mapa 713; Jacobs & Renner, 1974; Radisic, 1974; Harde, 1975:16; Athias, 1976:194; Nadolski, 1976; Andersson, 1977:100

Lucanus cervus (LINNAEUS); Clark, 1977:9

Leech, 1977:69; Perrier, 1977:167; Hempel & Schiemenz, 1978; Lacroix, 1978:253; Cameron, 1980:31; De Walsche, 1980:201; Krikken, 1980:24

Lucanus cervus (LINNAEUS); Roer, 1980:248-251

Numhauser, 1981:74; Boosten, 1982:96; Decelle, 1982:203; Español & Belles, 1982:73; Klausnitzer, 1982:14, 17; Bessonnat, 1984:77-80; Lojtnant, 1984; Fellenberg, 1985:79-80; Stolk, 1985:159; Bartolozzi, 1986:12; Cruysbergh, 1986:103; Chalmers-Hunt, 1987:184; Creutz, 1987:58-59; Long, 1987:323-324; Marples, 1987:149-150; Newton, 1987:371; Sakaino, 1987:16; Alexay, 1988:24-25; Andersson, Coulianos, Ehnstrom, Hammarsted, Imby, Janzon, Lindelow & Walden, 1987:70; Gjurasin, 1988:60-61; Grayson, 1988:224; Rabil, 1988:217; Verdcourt, 1988:36; Wootton, 1988:113; Bartolozzi, 1989:104; Carriere, 1989:19-20; George, 1989:28; Pfaff, 1989:31-32; Baker, 1990:12; Hein, 1990:34; Verdcourt, 1990:196

Lucanus cervus (LINNAEUS); Zemp, Hartmann, Knecht & Sprecher, 1990.

Bowdrey, 1991:300; Gouranton, Folliot & Thomas, 1991:102-109; Kranjcev, 1991:18-19; Kuhnel, 1991:209; Mamonov, 1991:157-163; Morris, 1991:106; Newton, 1991:36-38; Shepard & Biggs, 1991:7-8; Delatour, 1992:75; Alvarez-Lao, 1992:316-323; Maes, 1992:16-17; Spooner, 1992:329-330; Baraud, 1993:61; Bartolozzi & Onore, 1993:126

Lucanus cervus (LINNAEUS); Ochse, 1993:47-52.

Whitehead, 1993:206; Wuest, 1993:132; Hecq, 1994:455; Karner, 1994:71-72; Mizunuma & Nagai, 1994:212, pl. 8-9

Lucanus cervus (LINNAEUS); Klausnitzer, 1995.

Viejo & Sanchez Cumplido, 1995:186; Bartolozzi, 1996:486; Teunissen & Smit, 1996:134-135; Araya, 1996:19; Köteles & Bokony, 1996:99; Kral & Vitner, 1996:425

Lucanus cervus (LINNAEUS); López-Colón, González Peña & Beltrán Valen, 1996:16, 18.

Lucanus cervus cervus (LINNAEUS); López-Colón, González Peña & Beltrán Valen, 1996: 17.

Cecchi & Bartolozzi, 1997:122; Heynderycx, 1997:392-393, 396; Hoekstra, 1997:93-95.

Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 1998:142-166.

Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 1998:471-479.

Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 2001:2-11.

Lucanus cervus LINNAEUS; Sprecher-Uebersax & Durrer, 2001:161-182.

Lucanus cervus (LINNAEUS); Reyes-Castillo, 2002:486.

Lucanus cervus (LINNAEUS); Ballerio, 2003:68, 73, 82.

Norway

Sweden

Finland

Denmark

Great Britain : Borders (Eccles--55°37N-2°22W), Northumberland (Horsley--54°59N-1°09W), Cumbria (Keswick--54°36N-3°07W), W. Yorks (Burley--53°55N-1°41W, Apperley--53°51N-1°42W), Lancashire (Thornton--53°53N-3°01W), Clwyd (Prestatyn--53°20N-3°23W), Lincoln (Lincolnshire--53°13N-0°31W, Boston--52°59N-0°00--Billups), Nottingham (Nottingham--52°58N-1°09W), Derby (Derbyshire--52°56N-1°28W, Calke, Calke--Garneys), Shropshire (Wellington--52°40N-2°30W {Ercall wood, Apley wood, Ollerton wood}, Church Stretton--52°33N-2°49W), Hereford & Worcester (Bewdley--52°22N-2°20W, Bewdley--52°22N-2°20W--Blatch, Worcestershire--52°11N-2°14W), Warwick (Warwick--52°18N-1°34W, Warwick--52°18N-1°34W--Martineau), Cambridge (Cambridgeshire--52°12N-0°08E--Bloomfield), Buckingham (Buckinghamshire--52°01N-1°00W), Suffolk (Woodbridge--52°06N-1°18E, Ipswich--52°04N-1°09E, Nayland--51°58N-0°52E, Felixstowe--51°58N-1°20E), Essex (Halstead--51°57N-0°21E, Manningtree--51°56N-1°04E, Colchester--51°54N-0°54E, Braintree--51°53N-0°33E, Coggeshall--51°52N-0°41E, Marks Tey--51°52N-0°48E, Brightlingsea--51°49N-1°01E, Witham--51°48N-0°39E, West Mersea--51°47N-0°55E, Epping Forest--51°42N-0°06E, Brentwood--51°37N-0°18E, Tilbury--51°27N-0°22E), Gloucester (Cheltenham--51°54N-205°W, Birdlip--51°49N-2°07W, Newnham--51°48N-2°28W, Hardwicke--51°48N-2°18W), Hertford (Hatfield--51°47N-0°14W, Waltham Cross--51°42N-0°11W), Oxford (Oxford--51°46N-1°15W, Wallingford--51°36N-1°09W, Mapledurham Park), London (Enfield--           51°39N-0°05W, Pinner--51°36N-0°24W, Harrow--51°35N-0°21W = Harrow North, Ilford--51°34N-0°08E, Romford--51°34N-0°11E, Uxbridge--51°33N-0°29W, Upminster--51°33N-0°15E, London--51°32N-0°15W, Southall--51°31N-0°23W, Dagenham--51°31N-0°01E, Woolwich--51°29N-0°03E, Greenwich--51°28N-0°01E, Eltham--51°27N-0°03E, Hounslow--51°25N-0°21W, Hampton--51°25N-0°21W = Hampton Court, Kingston--51°25N-0°18W, Streatham--51°25N-0°08W, Bromley--51°25N-0°01E, Sidcup--51°25N-0°07E,       Wimbledon--51°23N-0°12W, Croydon--51°22N-0°06W, Sutton--51°21N-0°11W, Purley--51°20N-0°07W, Hatton Garden, Cadogan Square, New Forest), Glamorgan (Swansea--51°37N-3°56W), Buckingham (Marlow--51°35N-0°46W), Berkshire (Cookham--51°34N-0°42W, Maidenhead--51°31N-0°43W {Kimbers--Spring Hill}, Slough--            51°31N-0°36W, Pangbourne--51°29N-1°07W {Tidmarsh}, Windsor Forest--51°29N-0°37W, Datchet--51°29N-0°35W, Caversham--        51°28N-0°59W = Caversham Warren, Reading--51°27N-0°58W {Tilehurst}), Kent (Dartford--51°26N-0°14E, Gravesend--51°26N-0°22E, Rochester--51°25N-0°35E, Sheppey I.--51°25N-0°50E {Sheerness--51°26N-0°46E}), Surrey (Staines--51°26N-0°29W, Chertsey--51°24N-0°29W, Surbiton--51°24N-0°19W, Weybridge--51°23N-0°35W, Walton--51°23N-0°35W, Bagshot--51°22N-0°42W = Bagshot Heat, Epsom--51°20N-0°17W, Woking--51°19N-0°25W, Leatherhead--51°18N-0°20W, Box Hill--51°16N-0°20W, Merrow--51°15N-0°32W, Godalming--51°12N-0°37W, Chiddingford--51°07N-0°39W, Merton Park, Kew, Great Leight Rectory), Hampshire (Farnborough--51°17N-0°46W, Winchester--51°05N-1°19W, Bitterne--50°55N-1°22W, Southampton--50°55N-1°24W, Havant--50°52N-0°59W, New Forest--50°51N-1°35W, Beaulieu Abbey--50°49N-1°28W, Gosport--50°48N-1°08W, Lymington--50°46N-1°33W, Southsea--50°46N-1°04W), Somerset (Castle Cary--51°06N-2°31W), Wilts (Salisbury--51°05N-1°48W), Devon (Instow--51°03N-4°10W, Merton--50°53N-4°06W = Merton Park), W. Sussex (Midhurst--50°59N-0°45W, Arundel--50°51N-0°35W, Chichester--50°50N-0°48W, Shoreham-by-See--50°50N-0°17W = Shoreham, Worthing--50°49N-0°22W, West Tarring), E. Sussex (Brighton--50°46N-0°08W), Dorset (Lyme Regis--50°43N-2°57W), Cornwall (Yelverton--50°29N-4°05W, Saltash--50°24N-4°12W, Polperro--50°19N-4°31W, Truro--50°14N-5°03W), Middlesex (Teddington--Fulwell Vicarage, Heston, Lampton), Redmarley, Little Tring, Puttenham, Appleslade, Ellingham, Highwood, Linwood, Calmore, Hamble, Brightwell, Purbrook, Baughurst, Tadley, Mapledurham, Tidmarsh, Ewelme, Swyncombe Down, Bray, Henley, Eashing, Frimley, Wellington College, Brookwood, Sunninghill, Stoke Podges, Buryan, Fulwell, Mount Edgcumb, Bere Alston, Buckland Monachorum, Fingle Gorge, Slapton, Colyton, Parkestone, West Howe, Mudeford, Bransgore, West Cowes, Alverstoke, Kirby Bedon, Broxbourne, Chesnut, Wormley, Great Leighs, West Stow, Layer Breton, Layer-de-La-haye, Peldon, Tollesbury, Stanway, Boxted, Great Horkesley, Mount Bures, East Mersea, Fingringhoe, Arleigh, Elmstead, East Donyland, Thorpe-le-Soken, Wivenhoe, Dedham, Lawford, Thorrington Street, Kersey, Great Bentley, Little Bentley, Bradfield, Brantham, Holbrook, Woolverstone, Little Oakley, Guilford, Wisley, Oatlands, Shepperton, Northwood, Lancing, Sompting, Bramber, Henfield, Steyning, Brockham, Ashtead, Bookham, Ranmore, Chessington, Claygate, East Molesley, Esther, Hanworth, New Malden, Teddington, Fetcham, Heston, Kew, Lampton, Petersham, Richmond Park, Sheen, Carshalton, Mitcham, Barnes, Battersea, Cadogan Square, Chelsea, Chiswick, Fulham, Putney, Roehampton Lane, Wandsworth, Wandsworth Common, Addiscombe, Beckenham, Beddington, Norbury, Stoneleigh, Blackheat, West Dulwich, Bostall Heat, Brockley, Catford, Camberwell, Lambeth, Lee, Lewisham, South Norwood, Southwark, Sydenham, Hatton Garden, Holborn, Chingford, Barcombe, Abbey Wood, Charlton, Chislehurst, Danson Park, Plumstead Common, Upper Belvedere, Leyton, Loughton, Theydon Bois, Lullingstone, Darenth Park, Harold Wood, Larkfield, Grays, Stifford, West Thurrock, Aylesford, Birling, Ditton;

Netherlands : Drenthe, Overijssel--Twente, Gelderse Vallei, Utrecht (Utrechtse Heuvelrug), Noord-Brabant, Zuid-Limburg, Eysden, Alpesdroom;

Belgium : Anvers (Merxem, Postel), Limbourg (Lanaken--50°53N-5°39E, Vliermaalroot--50°50N-5°23E, Canne), Brabant (Louvain--50°52N-4°42E, Uccle--50°49N-4°21E, Overijse--50°46N-4°34E, Halle--50°44N-4°14E, Boistfort, Hoeilart, Bois de la Cambre, Etterbeek, Auderghem, Rouge-Cloître, Watermael, Boendael, Notre-Dame-au-Bois, Linkebeek, Rhode-Sainte-Genèse, Buysinghen, Assche), Liège (Wandre--50°40N-5°40E, Liège--50°38N-5°35E, Jupille--50°38N-5°38E, Chênée--50°37N-5°37E, Chaudfontaine--50°34N-5°40E, Huy--50°31N-5°15E, Sart--50°31N-5°57E, Spa--50°29N-5°52E, Aywaille--50°28N-5°40E, Bois d'Aywaille--50°28N-5°40E--Kin, Fléron, Ougrée, Embourg, Chèvremont, Beaufays, Tilff, Strée, Bois l'Evèque, Val Benoît, Jemeppe, Cointe, Beyne, Méry), Hainaut (Chimay--50°03N-4°19E--Forges-lez-Chimay, Harmignies, Vergnies), Namur (Namur--50°28N-4°52E, Hastière--50°14N-4°42E, Eprave, Foy-Notre-Dame, Waulsort, Marche-les-Dames), Luxembourg (Saint-Hubert--50°02N-5°23E, Bouillon--49°47N-5°05E, Laroche, Izel, Mardesous);

Luxembourg (Echternach--49°49N-6°26E--Berdorf);

France : Nord (Armentières--50°41N-2°52E), Calvados (Saint Aubain--49°20N-0°23W, Pont l'Evêque--49°17N-0°11E, Bures), Aisne (Vise), Marne (Chalons-sur-Marne--48°58N-4°22E), Meuse, Yvelines (Poissy--48°55N-2°05E, Saint Germain en Laye--48°52N-2°05E, Draveil, Jubisy, Bois de Boulogne, Auteuil, Longchamps, Achères, Neuilly, Vincennes, Vésinet, Croissy, Bougival, Marly), Val D'Oise (Montmorency--48°59N-2°22E), Hauts-De-Seine (Meudon--48°50N-2°15E), Paris (Paris--48°50N-2°20E), Essonne (Brétigny-sur-Orge--48°37N-2°18E), Finistère (Benodet--47°53N-4°07W), Morbihan (Grandchamp--47°46N-2°51E), Seine-Et-Marne (Melun--48°33N-2°38E--Mée), Terr. de Belfort (Delle--47°31N-6°59E), Indre-Et-Loire (Tours--47°25N-0°42E), Loir-Et-Cher (Romorantin--47°22N-1°44E), Cher (Meillant), Côte d'Or (Petigny), Allier (Saint Ennemond, Saint Evremond, Tronçais), Saône et Loire (Les Antoines, Issy l'Evêque), Charentes Maritimes (Montguyon--45°11N-0°12W), Charente (Féolle, Floirac), Dordogne, Limousin, Loire (Saint Etienne--45°28N-4°25E), Isère, Haute-Savoie (Chamonix--45°55N-6°51E, Annecy--45°54N-6°07E), Gironde (Bordeaux--44°50N-0°37W, La Ruscade), Lot-Et-Garonne (Agen--44°11N-0°40E), Tarn-Et-Garonne (Puylaroque, Montauban--44°00N-1°12E), Aveyron (Decazeville--44°33N-2°14E, Saint-George de Luzençon), Ardèche (Casteljau, Saint-Remèze, bois de Païolive), Drôme (Saint Donat sur l'Herbasse--45°07N-4°59E), Landes (Parentis--44°21N-1°07W, Dax--43°42N-1°02W), Gers (Escorneboeuf), Pyrénées-Atlantiques (Salies de Bearn--43°29N-0°58W, Saint Jean de Luz--43°21N-1°26W, Pau--43°18N-0°25W), Hautes Pyrénées (Lourdes--43°06N-0°02E, Cauterets--42°52N-0°03W, Lespoune), Ariège (Pamiers--43°06N-1°37E, Foix--42°03N1°36E), Tarn (Grésigne), Aude (Esperanza), Pyrénées Orientales (La Massane, Ria, Banyuls), Hérault (Montpellier--43°36N-3°57E, Saint Gély, Saint Guilhem, Minerve), Bouches du Rhône (Auriol--43°22N-5°38E, Marseille--43°17N-5°24E, Géménos--43°17N-5°37E, Saint Pons), Vaucluse (Mts. du Luberon--43°50N-5°20E, Oppéde), Alpes de Haute Provence (Banon--44°02N-5°37E, Riez--43°49N-6°05E), Var (Le Luc--43°24N-6°19E, La Roquebrussane--43°20N-5°59E, Mts. Sainte Baume--43°19N-5°45E, Forêt de Sainte-Baume--43°19N-5°45E, Pignans--43°18N-6°14E, Signes--43°17N-5°50E, Montrieux le Vieux--43°15N-5°58E, Collobrières--43°15N-6°19E, Belgentier--43°14N-6°01E, Toulon--43°07N-5°55E, Hyères--43°07N-6°04E, Correns, forêt de Morrières, Evenos, Aiguines, Mt. Touar, Valbelle), Alpes Maritimes (Saint Cézaire, Gilette), Corse--42°N-9°E, ? (Guyon, Saint Martin Vesubie);

Spain : Cantabria : Potes, 43°09 N - 4°37 W

Spain : Guipúzcoa : Tolosa, 43°09 N - 2°05 W

Spain : Cataluña

Spain : Huesca : Torla, 42°38 N - 0°06 W

Spain : Huesca : Benasque, 42°36 N - 0°31 W

Spain : Huesca : Valle de Gistain, 42°36 N - 0°20 E

Spain : Huesca : Plan, 42°35 N - 0°20 E

Spain : Huesca : Jaca, 42°34 N - 0°33 W

Spain : Huesca : Loarre, 42°19 N - 0°37 W

Spain : Huesca : La Peña

Spain : Teruel : Albarracín, 40°30 N - 1°34 W

Germany : Hamburg (Hamburg--53°32N-10°00E), Nordrhein-Westfalen (Dortmund--51°31N-7°27E), Hessen (Wetzlar--50°33N-8°31E, Kassel--51°19N-9°28E), Rheinland (Koblenz--50°22N-7°36E), Baden-Wurtemberg (Pforzheim--48°55N-8°42E), Rostock, Schwerin, Neubrandenburg, Magdeburg, Potsdam (Velten--52°41N-13°11E), Berlin, Frankfurt, Halle, Cottbus, Erfurt, Leipzig, Dresden, Suhl, Gera, Chemnitz = Karl-Marx-Stadt, ? (Kauden, Bentheumer Woud);

Poland;

Czech Rep. : Vinnéjezero, Pálava Biosphere Reserve;

Slovakia;

Switzerland : Basel : Kleinhuningen, 47°35 N – 7°35 E, 252 m, 1955 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Riehen, 47°35 N – 7°39 E : Schaferstrasse, 281 m, 1988 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Bachgraben, 275 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Baumlihof, 259 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Lange Erlen, 253 m, 1965 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Schwarz-Park, 261 m, 1992 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Bettingen, 47°34 N – 7°40 E : Chrischona, 470 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Basel : Basel, 47°33 N – 7°35 E (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Augst, 47°32 N – 7°43 E : Schlossli, 274 m, 1989 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Augst, 47°32 N – 7°43 E : Schonbuhl, 289 m, 1965 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Munchenstein, 47°31 N – 7°37 E : Hofmatt, 273 m, 1984 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Munchenstein, 47°31 N – 7°37 E : Ob. Wannen, 332 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Muttenz, 47°31 N – 7°39 E : Schanzli, 260 m, 1991 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Muttenz, 47°31 N – 7°39 E : Rutihard, 370 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Giebenacht, 47°31 – 7°48, 318 m, 1992 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Therwil, 47°30 N – 7°33 E : Froloo, 391 m, 1942 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Reinach, 47°30 N – 7°35 E : Leiwald, 310 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Arlesheim, 47°30 N – 7°37 E : Spitalholz, 348 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Arlesheim, 47°30 N – 7°37 E : Schwimmbad, 300 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Arlesheim, 47°30 N – 7°37 E : Reichenstein, 460 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Lausen, 47°29 N – 7°45 E, 344 m, 1984 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Blauen, 47°27 N – 7°31 E : Weekendhauser, 570 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Bubendorf, 47°27 N – 7°44 E : Wildenstein, 475 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Zwingen, 47°26 N – 7°32 E : Bahnhof, 339 m, 1995 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Himmelried, 47°26 N – 7°35 E: Steffen, 574 m, 1995 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Windentalhohe, 420 m, 1989 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Grammont, 539 m, 1989 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Schleifenberg, 534 m, 1990 (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Bern : Uetental, 380 m (citated by Sprecher-Uebersax & Durrer, 1998).

Switzerland : Jura : Lajoux, 47°17 N – 7°08 E

Switzerland : Vaud : Genève, 46°12N-6°09E

Switzerland : Bagatz

Osterreich : Ost-Tirol : Thuringen, 47°12 N - 9°46 E

Osterreich : Donau : Neuburg Bergheim

Osterreich : Wien, 48°12 N-16°21 E

Hungary : Mt. Bukk, 48°04N-20°29E

Hungary : Gödölló, 47°37N-18°54E

Hungary : Veszprém, 47°05N-17°57E

Hungary : Lake Balaton, 46°50N-17°45E : Tihany

Hungary : Mecsek Mts., 46°10N-17°56E

Hungary : Zebegewy

Hungary : Pomaz

Hungary : Borosjenö

Hungary : Torokbalint

Romania

Italy : Piemonte (Arona--45°45N-8°32E, Biella--45°34N-8°01E), Lombardia (Varese--45°49N-8°51E--Montvalle, Como--45°48N-9°05E), Venezia Giulia (Udine--46°05N-13°12E, Udine--46°05N-13°12E--Toppo, Vittorio--45°58N-12°17E, Pordenone--45°57N-12°38E, Treviso--45°39N-12°14E, Trieste--43°40N-13°46E {San Sergio-Petrigna, Valle Ostra}), Liguria (Genova--44°22N-8°55E), Toscana (Firenze--43°47N-11°16E = Florence, Siena--43°19N-11°19E, Cortona--43°15N-12°00E, Badia Prataglia), Emilia Romagna (La Lama), Sardegna--40°10N-9°E, Sicilia--37°35N-14°15E, ? (Zattaglia);

Slovenija: Mackovci--46°47N-16°10E, Maribor--46°34N-15°37E, Celje--46°15N-15°14E, Celje--46°15N-15°14E--Ljubno ob Savinja River, Ljubljana--46°04N-14°29E, Litija--46°03N-14°52E--Hotic, Grosuplje--45°57N-14°40E--Tabor, Ajdovscina--45°53N-13°53E, Cerknica--45°48N-14°20E, Postojna--45°47N-14°11E--Javorniki, Sezana--45°43N-13°50E, Kocevje--45°39N-14°49E, Kozina--45°34N-13°56E, Bohinj--Ribcev laz, Bohinj--Vogar, ob Suhi-Bohinj, Bohinjska Bistrica, Crna prst, Gameljne, Gorica, Kamnik, Komenda, Kostanjevica, Kuzma, Lubnik, Orehovci--Gornja Radgona, Pekel--Zalec, Podgorje--Slavnik, Radenci, Rakek, Rakov Skocjan, Senozece, Sorica, Sorsko polje, Tacen, Tatre, Trzin, Vaneca, Vipava, Vodice, Zg. Tuhinj;

Bosnia & Hercegovina : Hercegovina (Konjice--43°39N-17°56E, Mostar--43°20N-17°48E);

Yugoslavia : ? (Karlovak);

Bulgaria;

Greece : Karpenision--38°55N-21°46E;

Turkey;

Russia : to Ural and Caucasus (Terek River, Khosta);

Iran.

Note : this species is protected by Bern Convention appendix 3; Habitat Direction appendix 2; Regional Law of Toscana, Italia, appendix A and B (Ballerio, 2003).

Xylophagous :

Betulaceae : Alnus, Carpinus.

Fagaceae : Quercus, Fagus.

Hippocastanaceae : Aesculus.

Juglandaceae : Juglans.

Malaceae : Pyrus, Malus.

Moraceae : Morus.

Oleaceae : Fraxinus.

Pinaceae : Picea, Pinus.

Rosaceae : Cerasus, Prunus.

Salicaceae : Salix, Populus.

Tiliaceae : Tilia.

Ulmaceae : Ulmus.

Saproxylophagous :

Aceraceae : Acer.

Fagaceae : Quercus.

Can breed in compost (artificial soil).

Natural enemies :

Birds : Corvidae : Pica.

? : Caprimulgus europaeus, Strix aluco.

Acarina : Uropodidae : Polyaspis patavinus.

? : Isacis cuspidat, Isacis lucani, Megaselia rufipes, Monocercomonas melolonthae.

Bibliografía :

BAGUENA L. (1967) Scarabaeoidea de la fauna íbero-balear y pirenaica. C.S.F.C. Madrid, 1955 (publicado en 1967): 1-567.

BALLERIO A. (2003) EntomoLex : la conservazione degli insetti e la legge. Memorie Soc. entomol. Italiana, 82(1):17-86.

COFAIS M. (1972) Un cas tératologique chez Lucanus cervus. L’Entomologiste, 28:32-33.

CLARK J.T. (1977) Aspects of variation in the sag beetle. Systematic Entomology, 2:9-16.

FREUDE H. (1968) Uber Mandibel-Anomalitaten bei Lucanus cervus. Nachrichtenblatt der Bayerischen Entomologen, 17:59-64.

HORION A. (1958) Faunistik der mitteleuropaischen Kafer 6: Lamellicornia. 343 pp.

KLAUSNITZER B. (1995) Die Hirschkafer. Neue Brehm-Bucherei, 551:109 pp.

LOPEZ-COLON J.I., GONZALEZ PEÑA C.F. & BELTRAN VALEN J.R. (1996) Familia : Lucanidae. Catalogus de la Entomofauna Aragonesa, No. 12:15-20.

MAL N. (1972) Sur le polymorphisme antennaire d’une population provençale de Lucanus cervus. L’Entomologiste, 38:161-166.

OCHSE M. (1993) Angaben zum Hirschkafer, Lucanus cervus L., bei Kassel. Hessische faunistische Briefe, 13:47-52.

REYES-CASTILLO P. (2002) 27. Lucanidae. Pp. 485-492. En LLORENTE BOUSQUETS J. & MORRONE J.J. Biodiversidad, Taxonomía y Biogeografía de Artrópodos de México : Hacia una síntesis de su conocimiento. Universidad Nacional Autónoma de México. Vol. III:692 pp.

ROER H. (1980) Zur Verbreitung und Bestandesdichte des Hirschkafers im Rheinland. Acta Musei Rigenaehradecensis, Serie A, Supplement: 248-251.

SPRECHER-UEBERSAX E. & DURRER H. (1998) Uber das Vorkommen des Hirschkafers (Lucanus cervus L.) in der Region Basel (Coleoptera). Mitt. Ent. Ges. Basel, 48(4):142-166.

SPRECHER-UEBERSAX E. & DURRER H. (1998) Untersuchungen zum Stridulationsverhalten der Hirschkafer-Larven (Lucanus cervus L.) (Coleoptera : Lucanidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71:471-479.

SPRECHER-UEBERSAX E. & DURRER H. (2001) Beobachtungen zur Nahrungswahl des Hirschkafers (Lucanus cervus L.) (Coleoptera). Mitteilungen der Entomologischen Gesellschaft Basel, 51(1):2-11.

SPRECHER-UEBERSAX E. & DURRER H. (2001) Verhaltenstudien beim Hirschkafers mittels Telemetrie und Videoaufzeichnungen (Coleoptera, Lucanus cervus L.). Mitteilungen der Naturforschenden Gesellschaften beider Basel, 5:161-182.

ZEMP M., HARTMANN K., KNECHT D. & SPRECHER-UEBERSAX E. (1990) Arlesheimer Wald Spitalholz 1990. Unveroffentliches Manuskript zuhanden der Kober-Schwabe-Stiftung Arlesheim, 59 pp.

 

Lucanus (Lucanus) cervus ssp. fabiani MULSANT & GODART, 1855 [Platycerus].

Hexaphyllus pontbrianti MULSANT 1839 [Lucanus, Platycerus] [France : Lyon : Bois de Rochecardon].

Lucanus pentaphyllus REICHE 1853 [Platycerus] [Gallia meridionalia : circa Telonem et Portum Veneris].

Lucanus heptaphyllus ABEILLE DE PERRIN 1867.

Lucanus reichei MOTSCHULSKY 1870 [France : Toulouse].

Lucanus capreolus hexaphyllus (PLANET) DIDIER & SEGUY 1953 [Italie] (nomen nudum).

SE. France : Rhône (Lyon--45°45N-4°51E--bois de Rochecardon), Ardèche, Drôme (Nyons--44°21N-5°08E), Landes (Dax--43°42N-1°02W), Pyrénées Orientales (Prades--42°37N-2°25E, Port-Vendres--42°29N-3°07E, Ria, Vernet-les-Bains, Banyuls-sur-Mer), Hérault (Montpellier--43°36N-3°57E), Var (La Roquebrussane--43°20N-5°59E, Montrieux le Vieux--43°15N-5°58E, Toulon--43°07N-5°55E, Hyères--43°07N-6°04E), Alpes Maritimes, Corse (St. Hyacinthe, Ponte Vecchiu near Ota).

Note : For Lacroix (1968) fabiani is only a morfa of cervus because cervus and fabiani are living in the same region.

 

Lucanus (Lucanus) cervus ssp. mediadonta LACROIX, 1978.

Georgia : Tiflis*.

 

Lucanus (Lucanus) cervus ssp. tauricus MOTSCHULSKY, 1845.

Lucanus maxillaris MOTSCHULSKY 1845 [Crimée, Caucase, Tourcménie].

Lucanus brevicollis MOTSCHULSKY 1870 [SW European Russia : Voronège].

Lucanus europaeus MOTSCHULSKY 1870 [Kharcov].

Russia : Voronesk;

Ucrania : Kharkov, Krym = Crimea*;

Turkmenistan.

 

Lucanus (Lucanus) cervus ssp. judaicus PLANET 1902.

Turkey : Aïn-Tab* at 90 km NE Alep.

 

Lucanus (Lucanus) cervus ssp. turcicus STURM, 1843 [Platycerus].

Romania : Bucaresti--44°25N-26°07E--Ilfov;

Bulgaria : Balchik--43°24N-28°09E, Arkutino, Lozenec, Skorpilovce;

Greece : Eubée I.--38°35N-23°50E;

Turkey : Istanbul--41°02N-28°56E = Constantinople, Istambul--41°02N-28°56E--Forest de Belgrad, Bursa--40°14N-29°01E, Taurus--Adana--36°56N-35°20E, Alyana.

 

Lucanus (Lucanus) cervus ssp. akbesianus PLANET, 1896.

Lucanus cervus turcicus ab. syriacus MOTSCHULSKY (nec MOTSCHULSKY) 1897.

Lucanus cervus ab. motschulskyi BENESH 1960.

Lucanus poujadei PLANET 1897 [Syrie].

Lucanus smyrniacus DIDIER & SEGUY 1953 [Asie mineure].

Lebanon : Djezin;

Syria : Akbes* = Sgée, Marah, Smyrne, Alexandrette, Ghazir, Marsch.

 

Lucanus (Lucanus) convexus LACROIX, 1982.

Lacroix, 1982:14; Maes, 1992:17.

India : Nagaland*.

 

Lucanus (Lucanus) cyclommatoides DIDIER, 1928.

Lucanus cyclommatoides DIDIER, 1928:51

Lucanus cyclommatoides DIDIER; Didier & Séguy, 1953:81

Lucanus cyclommatoides DIDIER; Benesh, 1960:140

Lucanus cyclommatoides DIDIER; Maes, 1992:17.

Lucanus cyclommatoides DIDIER; Araya, 2001:12, figs.16-20.

·        Vietnam : Tonkin : Chapa, 13-VII, col. Jeanvoine (male TYPE of Lucanus cyclommatoides DIDIER, 1928 en Museum National d’Histoire Naturelle, Paris).

·        Vietnam : Tonkin : Chapa, 26-IX-1918, col. Jeanvoine, ex. coll. Boileau (female COTYPE of Lucanus cyclommatoides DIDIER, 1928 en Museum National d’Histoire Naturelle, Paris).

Male TYPE of Lucanus cyclommatoides DIDIER, 1928, from Vietnam : Chapa (adapted from Araya, 2001)

Female COTYPE of Lucanus cyclommatoides DIDIER, 1928, from Vietnam : Chapa (adapted from Araya, 2001)

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) delavayi FAIRMAIRE, 1887.

Lucanus delavayi FAIRMAIRE, 1887:27 [China : Yunnan].

Lucanus delavayi FAIRMAIRE; Fairmaire, 1887:98

Lucanus delavayi FAIRMAIRE; Planet, 1900:228

Lucanus delavayi FAIRMAIRE; Planet, 1902:68

Lucanus delavayi FAIRMAIRE; Heyne & Taschenberg, 1908:50

Lucanus delavayi FAIRMAIRE; Didier, 1949:pl.10

Lucanus delavayi FAIRMAIRE; Benesh, 1960:140

Lucanus delavayi FAIRMAIRE; Kurosawa, 1978:143

Lucanus delavayi FAIRMAIRE; Bomans, 1989:9

Lucanus delavayi FAIRMAIRE; Maes, 1992:17

Lucanus delavayi FAIRMAIRE; Mizunuma & Nagai, 1994:216, pl.16.

Lucanus delavayi FAIRMAIRE; Araya, 2001:13, figs.25-26.

·        China : Yunnan, col. R.P. Delavay (male PARATYPE of Lucanus delavayi FAIRMAIRE, 1887, # 1666 in Museum d’Histoire Naturelle, Geneva).

·        China : Yunnan : Ta-Pin-Tze

·        China : Yunnan : Yunnan-Sen

·        China : Yunnan : San-Nen-Kai.

male PARATYPE of Lucanus delavayi FAIRMAIRE, 1887, from China : Yunnan (adapted from Araya, 2001)

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) deuveianus BOUCHER, 1998.

Lucanus deuveianus BOUCHER, 1998:182-184, fig. 2, 4 [China : Guangxi : Daming Shan].

Lucanus deuveianus BOUCHER; Araya, 2001:14, figs.29-30.

·        China : Guangxi : Daming Shan, 1200 m, VI-1995, col. T. Deuve (male HOLOTYPUS of Lucanus deuveianus BOUCHER, 1998 in Museum National d'Histoire Naturelle, Paris).

male HOLOTYPUS of Lucanus deuveianus BOUCHER, 1998, from China : Guangxi : Daming Shan (adapted from Araya, 2001).

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) didieri PLANET, 1927.

Lucanus didieri PLANET, 1927:106 [China : Yunnan : Pe-Yen-Tsin]

Lucanus didieri PLANET; Didier & Séguy, 1952:pl.15

Lucanus didieri PLANET; Benesh, 1960:140

Lucanus didieri PLANET; Lacroix & Bomans, 1973:145

Lucanus didieri PLANET; Maes, 1992:17

Lucanus didieri PLANET; Mizunuma & Nagai, 1994:309, pl.147.

Lucanus didieri PLANET; Araya, 2001:14, figs.31-32.

·        China : Yunnan : Pe-Yen-Tsin, ex. col. Le Moult (male TYPUS of Lucanus didieri PLANET, 1927 in Museum National d'Histoire Naturelle, Paris).

male TYPUS of Lucanus didieri PLANET, 1927, from China : Yunnan : Pe-Yen-Tsin (adapted from Araya, 2001).

Bibliography :

ARAYA K. (2001) Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.

 

Lucanus (Lucanus) dohertyi BOILEAU, 1911.

Lucanus laminifer var. minor WATERHOUSE 1890.

Waterhouse, 1890:33; Boileau, 1911:435, pl.34; Arrow, 1950:56; Didier & Séguy, 1952:pl.15; Benesh, 1960:140; Lacroix & Bomans, 1973:138, 142; Lacroix, 1982:13; Téllez, 1990:39; Maes, 1992:17; Mizunuma & Nagai, 1994:214, 309, pl.11.

India : Assam, Meghalaya (Khasi Hills), Nagaland (Naga Hills*).

 

Lucanus (Lucanus) elaphus FABRICIUS, 1775.

Lucanus cervus var. DEGEER 1774:33.

Lucanus elaphus FABRICIUS, 1775:2 [Virginia].

Lucanus elaphus FABRICIUS; Fabricius, 1781:2.

Lucanus elaphus FABRICIUS; Fabricius, 1887:1.

Lucanus elaphus FABRICIUS; Olivier, 1789:12-13, pl.3.

Lucanus elaphus FABRICIUS; Linnaeus (ed. Gmelin), 1790:1589.

Lucanus elaphus FABRICIUS; Fabricius, 1792:237.

Lucanus elaphus FABRICIUS; Olivier (ed. Illiger), 1800:60-61.

Lucanus elaphus FABRICIUS; Fabricius, 1801:249.

Lucanus elaphus FABRICIUS; Olivier (ed. Sturm), 1802:2, pl.2.

Lucanus elaphus FABRICIUS; Thunberg, 1806:191.

Lucanus elaphus FABRICIUS; Schonherr, 1817:322.

Lucanus elaphus FABRICIUS; Dejean, 1837:193.

Lucanus elaphus FABRICIUS; Burmeister, 1847:354.

Lucanus elaphus FABRICIUS; Fuchs, 1882:49.

Lucanus elaphus FABRICIUS; Albers, 1890:88.

Lucanus elaphus FABRICIUS; Planet, 1902:92, pl.16.

Lucanus elaphus FABRICIUS; Ulke, 1902.

Lucanus elaphus FABRICIUS; Wickham, 1903:205-206.

Lucanus elaphus FABRICIUS; Heyne & Taschenberg, 1908:50, pl.6.

Lucanus elaphus FABRICIUS; Blatchley, 1910:905.

Lucanus elaphus ab. carlengi ANGELL 1916:70 [Kentucky : Louisville].

Lucanus elaphus FABRICIUS; Brimley, 1938.

Lucanus elaphus FABRICIUS; Loding, 1945.

Lucanus elaphus FABRICIUS; Didier, 1949:pl.11.

Lucanus elaphus FABRICIUS; Benesh, 1960:140-141.

Lucanus elaphus FABRICIUS; Arnett, 1965.

Lucanus elaphus FABRICIUS; Ritcher, 1966.

Lucanus elaphus FABRICIUS; Kirk, 1969.

Lucanus elaphus FABRICIUS; Kirk, 1970.

Lucanus elaphus FABRICIUS; Arnold & Drew, 1987.

Lucanus elaphus FABRICIUS; McNamara, 1991.

Lucanus elaphus FABRICIUS; Ratcliffe, 1991:267.

Lucanus elaphus FABRICIUS; Maes, 1992:17.

Lucanus elaphus FABRICIUS; Anonymous, 1994.

Lucanus elaphus FABRICIUS; Mizunuma & Nagai, 1994:212, pl. 7.

Lucanus elaphus FABRICIUS; Staines, 1994:17-18.

Lucanus elaphus FABRICIUS; Poole & Gentili, 1996:297.

Lucanus elaphus FABRICIUS; Staines, 2000:26-30.

Lucanus elaphus FABRICIUS; Staines, 2001:397-404.

Canada :

Ontario (citated by Benesh, 1960).

USA :

Minnesota (National Museum of Natural History, Washington).

Michigan (Ohio State University).

Nebraska : Richardson Co. : Stella, 6-VI-1999 (University of Nebraska).

Iowa : Lee Co. : Keokuk (Museum of Comparative Zoology, Cambridge).

Kansas (Field Museum of Natural History, Museum of Comparative Zoology, Cambridge, National Museum of Natural History, Washington).

Kansas : Douglas Co. (University of Kansas).

Kansas : Labette Co. : Oswego (National Museum of Natural History, Washington).

Kansas : Leavenworth Co. : Leavenworth, 23-VII-1943 (California Academy of Science).

Kansas : Leavenworth Co. : Leavenworth, 30-VII-1947 (University of Kansas).

Kansas : Wilson Co. : New Albany (Natural History Museum, London).

Kansas : Wyandotte Co. (Field Museum of Natural History).

Missouri (Field Museum of Natural History).

Missouri : Boone Co. : Columbia, VI-1903 (Field Museum of Natural History).

Missouri : Boone Co. : Columbia, 21-VI-1971 (University of Missouri).

Missouri : Boone Co. : Columbia, 14-VII-1959 (University of Missouri).